Flora of New York/Lycophytes

Lycophytes Dipsacales Ferns 1 Pre-polypodferns Order Lycopodiales The Lycopodiales ... Family Lycopodiaceae The Lycopodiaceae (clubmoss family) can, for identification purposes in New York State, roughly be divided into clumping (firmoss) and creeping (clubmoss) groups. 19 2 clumping(firmoss) #baa 12 creeping(clubmoss) #aba Lycopodiaceae Huperzioideae (firmoss) Huperzia lucidula (shining firmoss) #baa Huperzia appressa (mountain firmoss) #baa Huperzia selago (northern firmoss) #baa Huperzia × bartleyi (Bartley’s firmoss) #baa Lycopodielloideae bog clubmoss Lycopodiella inundata (northern bog-clubmoss) #aba Lycopodiella appressa (southern bog-clubmoss) #aba Lycopodiella alopecuroides (foxtail bog-clubmoss) #aba slender bog-clubmoss Pseudolycopodiella caroliniana (Carolina clubmoss) #aba Lycopodioideae running clubmoss Lycopodium clavatum (staghorn clubmoss) #aba Lycopodium lagopus (one-cone clubmoss) #aba tree clubmoss Dendrolycopodium obscurum (flat-branched tree clubmoss) #aba Dendrolycopodium dendroideum (prickly tree clubmoss) #aba Dendrolycopodium hickeyi (Hickey’s tree clubmoss) #aba bristly clubmoss Spinulum annotinum (common bristly clubmoss) #aba Spinulum canadense (northern bristly clubmoss) #aba groundcedar Diphasiastrum digitatum (southern ground cedar) Diphasiastrum tristachyum (blue ground cedar) Diphasiastrum complanatum (northern ground cedar) Diphasiastrum sitchense (Sitka ground cedar) #aba Subfamily Huperzioideae The Huperzioideae, known as firmosses, have also been treated as a separate family Huperziaceae. Firmosses differ from other Lycopodiaceae in that they grow in clumps, instead of producing creeping, horizontal stems like other family members."Family: Huperziaceae - firmoss family" Go Botany 3.5, Native Plant Trust, 2021 Of the three genera in this group, Huperzia is a temperate and arctic genus and is the only one found naturally in New York. Of the other two genera, Phlegmariurus is native to tropical and subtropical regions, and Phylloglossum is native to Australia and New Zealand. Huperzia Shining firmoss.jpg Huperzia lucidula, shining firmoss Commonly known as firmoss or fir-clubmoss, Huperzia (hoo-per-zee-uh) was named for Johann Peter Huperz, an early 19th century German botanist who specialized in ferns. The rare Appalachian firmoss was published as Huperzia appalachiana in 1992, but authorities, including Weakly (2023)Weakley, A.S., and Southeastern Flora Team. 2023. Flora of the southeastern United States. University of North Carolina Herbarium, North Carolina Botanical Garden, Chapel Hill, U.S.A. Dec 23, 2023. consider it to be indistinct from the previously published Huperzia appressa. Lycopodiales Lycopodiaceae Huperzioideae Huperzia Huperzia Firmoss 735 4 Bernh. (1801) Huperzia lucidula (Michx.) Trevis. (Michaux) Trevisan Atti Soc. Ital. Sci. Nat. 17: 248. (1875) 1803 Lycopodium lucidulum Michx. 1875 Huperzia lucidula (Michx.) Trevis. 1903 Huperzia selago (L.) Bernh. ex Schrank & Mart. lucidulum Clute 1937 Lycopodium lucidulum Michx. verum Wherry 1939 Urostachys lucidulus (Michx.) Herter ex Nessel 1939 Urostachys lucidulus (Michx.) Herter ex Nessel reclinatus (Dup) Nessel 1965 Huperzia selago (L.) Bernh. ex Schrank & Mart. lucidula (Michx.) Á.Löve & D.Löve 1967 Lycopodium lucidulum Michx. tryonii Mohlenbr. 1999 Huperzia lucidula (Michx.) Trevis. tryonii (Mohlenbr.) Mohlenbr. Shining firmoss Shining clubmoss Shining fir-clubmoss Lycopode brillant Huperzie brillant Native Secure 5 FAC FACW Perennial Subshrub S5, G5 1888 5 HULU2 NN 82407-Huperzia-lucidula 124855-2 2 6585 huperzia/lucidula 503079 456039 200002697 Huperzia_lucidula 26609538 Huperzia+lucidula 124855-2 Huperzia lucidula nymap.svg Lycopodium lucidulum témis 2x3.jpg Huperzia appressa (Bach.Pyl. ex Desv.) Á.Löve & D.Löve Bot. Not. 114: 34 (1961) 1827 Lycopodium selago L. appressum Bach.Pyl. ex Desv. 1902 Lycopodium selago L. appressum (Bach.Pyl. ex Desv.) Gelert 1930 Lycopodium appressum (Bach.Pyl. ex Desv.) Petrov Fl. Iakut. 1: 37 (1930), nom. illeg. 1939 Urostachys selago (L.) Herter appressus (Bach.Pyl. ex Desv.) Herter ex Nessel Bärlappgewächse: 38 (1939) 1956 Lycopodium selago L. appressa (Bach.Pyl. ex Desv.) D.Löve 1961 Huperzia appressa (Bach.Pyl. ex Desv.) Á.Löve & D.Löve Bot. Not. 114: 34 (1961) 1968 Lycopodium selago L. appressum (Bach.Pyl. ex Desv.) Hultén 1981 Huperzia selago (L.) Bernh. ex Schrank & Mart. appressa (Bach.Pyl. ex Desv.) Ching 1992 Huperzia appalachiana Beitel & Mickel Amer. Fern J. 82: 45 (1992) Appalachian firmoss Appalachian clubmoss Appalachian fir-clubmoss Mountain firmoss Mountain fir-clubmoss Lycopode des Appalaches Native Rare 9 3appalachian-firmoss Huperzia appressa Rare S3 G5 FACU Perennial Subshrub S3, G4-G5 1886 3 HUAP NN as Huperzia appalachiana Beitel & Mickel BCXPJ 3992299 204155-Huperzia-appressa 124840-2 0001274871 1 6583 huperzia/appressa 507547 233500681 Huperzia_appalachiana Huperzia+appressa 17440620-1 Huperzia appressa nymap.svg Essex, Greene Essex County Huperzia appressa 2x3.jpg Huperzia selago (L.) Bernh. ex Schrank & Mart. Hort. Reg. Monac.: 3 (1829) 1753 Lycopodium selago L. 1779 Lycopodium densum Lam. (nom. superfl.) 1805 Plananthus selago (L.) P.Beauv. 1819 Plananthus fastigiatus Opiz 1821 Lycopodium abietiforme Gray 1829 Huperzia selago (L.) Bernh. ex Schrank & Mart. 1831 Lycopodium selago L. densum Wallr. 1866 Selago vulgaris Schur 1923 Urostachys selago (L.) Herter 1942 Lycopodium selago L. typicum Wherry (not validly publ.) 1955 Mirmau selago H.P.Fuchs 1997 Huperzia acuminata (Sugim.) Satou Northern fir-moss Fir clubmoss Lycopode sélagine Huperzie sélagine Native Endangered 10 1fir-clubmoss Huperzia selago Endangered S1 G5 1883 1 HUSE NN as var. selago 119147-Huperzia-selago 66388 6588 huperzia/selago 200002709 456041 200002709 26603630 Huperzia+selago 1140250-2 Huperzia selago nymap.svg 5 counties 2 counties Huperzia selago 240906 2x3.jpg Huperzia porophila (F.E.Lloyd & Underw.) Holub 1900 Lycopodium porophilum F.E.Lloyd & Underw. 1903 Lycopodium selago (L.) Bernh. ex Schrank & Mart. porophilum Clute 1905 Lycopodium lucidulum Michx. porophilum (F.E.Lloyd & Underw.) Clute 1908 Lycopodium lucidulum Michx. porophilum (F.E.Lloyd & Underw.) Clute 1923 Urostachys porophilus (F.E.Lloyd & Underw.) Herter 1939 Urostachys lucidulus (Michx.) Herter ex Nessel porophilus (F.E.Lloyd & Underw.) Nessel 1965 Huperzia selago (L.) Bernh. ex Schrank & Mart. porophila (F.E.Lloyd & Underw.) Á.Löve & D.Löve 1985 Huperzia porophila (F.E.Lloyd & Underw.) Holub Rock clubmoss Rock firmoss Lloyd's clubmoss Lycopode porophile N. America native Excluded X 129 HUPO2 NN 163858-Huperzia-porophila 278425-2 3 17189 233500684 26610804 Excluded nymap.svg Excluded Carbon County, PA Huperzia porophila-2a.jpg Huperzia occidentalis (Clute) Kartesz & Gandhi Phytologia 70: 201. (1991) 1903 Lycopodium lucidulum Michx. occidentale Clute 1903 Lycopodium lucidulum Michx. occidentalis Clute 1932 Lycopodium lucidulum Michx. occidentale (Clute) L.R.Wilson 1966 Lycopodium selago L. occidentale (Clute) B.Boivin 1966 Lycopodium selago L. occidentale (Clute) B.Boivin 1967 Lycopodium lucidulum Michx. tryonii Mohlenbr. 1991 Huperzia occidentalis (Clute) Kartesz & Gandhi 1992 Huperzia occidentalis (Clute) Beitel (nom. illeg.) Pacific Clubmoss 2017 NatureServe Western Clubmoss 2004-2023 Burke Museum Herbarium, Seattle, WA Lycopode de l'Ouest 2017 NatureServe N. America native Northwest Excluded Perennial Herb X 129 EXCLUDED HUOC NN 2688528 143696-Huperzia-occidentalis 278409-2 503081 233500683 50052154 Huperzia+occidentalis Huperzia%20occidentalis Excluded nymap.svg Excluded Northwest N.A. only Huperzia occidentalis iNat-256345356.jpg Huperzia hybrids Lycopodiales Lycopodiaceae Huperzioideae Huperzia Huperzia Firmoss 735 4 Bernh. (1801) Huperzia × bartleyi (Cusick) Holub Folia Geobot. Phytotax. 26: 92 (1991) Huperzia lucidula Huperzia porophila 1987 Lycopodium × bartleyi Cusick 1991 Huperzia × bartleyi (Cusick) Holub 1991 Huperzia × bartleyi (Cusick) Kartesz & Gandhi Bartley's clubmoss Native Unranked 1870 U HUBA N0 AR, IA, IL, KY, MO, NC, NY, OH, SC, WI 170791-Huperzia-×-bartleyi 17543650-1 503069 50241448 Huperzia × bartleyi nymap.svg Ulster County No NY obsevations Ny hybrid.svg Huperzia × buttersii (Abbe) Kartesz & Gandhi Huperzia lucidula Huperzia selago 1953 Lycopodium × buttersii Abbe 1991 Huperzia × buttersii Kartesz & Gandhi Butters' club-moss Butters' firmoss Lycopode de Butters N. America native Excluded X 633 HUBU NN 170792-Huperzia-×-buttersii 278330-2 6589 huperzia/lucidula 503070 200002697 50241451 Huperzia+x+buttersii Excluded nymap.svg Ny hybrid.svg Huperzia × protoporophila A.Haines Fam. Huperziac. Lycopodiac. New England: 88 (2003) Huperzia appressa Huperzia lucidula 2003 Huperzia × protoporophila A.Haines Hybrid rock firmoss N. America native No N.Y. reports 0 D 170796-Huperzia-×-protoporophila 6591 huperzia/appressa 50305405 60453031-2 Excluded nymap.svg Ny hybrid.svg Huperzia × josephbeitelii A.Haines Fam. Huperziac. Lycopodiac. New England, ed. 2: 100 (2003) Huperzia appressa Huperzia selago 2003 Huperzia × beitelii A.Haines 2003 Huperzia × josephbeitelii A.Haines Beitel's firmoss Joseph Beitel's clubmoss N. America native No N.Y. reports 0 D HUJO NN north of NY, not in NY 170795-Huperzia-×-josephbeitelii 6590 huperzia/appressa 50305406 60453016-2 Excluded nymap.svg Ny hybrid.svg Subfamily Lycopodioideae Lycopodium 200px|thumb|right|Running clubmoss The genus name Lycopodium (lī-kə-ˈpō-dē-əm) was derived from the Greek lukos (wolf) and podion (small foot). It is commonly known as running clubmoss. Also treated as Lycopodium sect. Lycopodium. Lycopodiales Lycopodiaceae Lycopodioideae Lycopodium Lycopodium Running clubmoss 737 2 Lycopodium clavatum L. Sp. Pl.: 1101 (1753) 1753 Lycopodium clavatum L. 1805 Lepidotis clavata (L.) P.Beauv. 1805 Lepidotis ciliata P.Beauv. 1805 Lepidotis inflexa P.Beauv. 1810 Lepidotis aristatum Humb. & Bonpl. ex Willd. 1925 Lycopodium clavatum L. laurentianum Vict. 1925 Lycopodium clavatum L. subremotum Vict. Common clubmoss Running clubmoss Running pine Running cedar Staghorn clubmoss Elk moss Lycopode à massue Native Secure 5 1887 5 LYCL NN 304307-2 lycopodium/clavatum 77914-Lycopodium-clavatum 17024 431024 200002751 Lycopodium_clavatum Lycopodium clavatum nymap.svg LycopodiumClavatum.jpg Lycopodium lagopus (Laest. ex Hartm.) Zinserl. ex Kuzen 1831 Lycopodium clavatum L. monostachyon Hook. & Grev. 1838 Lycopodium clavatum L. integrifolium (Hook. ex Goldie) Spring 1842 Lycopodium clavatum L. integerrimum Spring 1858 Lycopodium clavatum L. lagopus Laest. ex C.Hartm. 1901 Lycopodium clavatum L. brevispicatum Peck 1910 Lycopodium clavatum L. megastachyon Fernald & Bissell 1925 Lycopodium clavatum L. furcatum Vict. 1950 Lycopodium clavatum L. monostachyon (Hook. & Grev.) Selander 1953 Lycopodium lagopus (Laest. ex C.Hartm.) Zinserl. ex Kuzen 1958 Lycopodium clavatum L. megastachyon (Fernald & Bissell) Á.Löve & D.Löve One-cone clubmoss One-cone ground-pine Running pine Running cedar Arctic stag's-horn clubmoss Lycopode patte-de-lapin Native Likely secure 4 1879 4 LYLA5 NN 1159641-2 lycopodium/lagopus 82390-Lycopodium-lagopus 507548 233500762 Lycopodium_lagopus Lycopodium lagopus nymap.svg Lycopodium lagopus iNat-100397035.jpg Dendrolycopodium thumb|right|150px|Tree clubmoss From Greek δενδρος (denros: tree), λυκος (lukos: wolf), and ποδιων (podion: legs), Dendrolycopodium (den-drō-lī-kə-ˈpō-dē-əm) contains tree-clubmoss species, which are fairly erect and have branched stems. This genus, also treated as Lycopodium sect. Obscura, is characterized by stalked terminal Strobilus strobili, which contain their Sporangium sporangia. Lycopodiales Lycopodiaceae Lycopodioideae Dendrolycopodium Dendrolycopodium Tree clubmoss 1084 3 Dendrolycopodium obscurum (L.) A.Haines 1753 Lycopodium obscurum L. 1833 Lycopodium dendroideum Michx. obscurum (L.) Torr. ex Beck 1937 Lycopodium obscurum L. genuinum Wherry 2003 Dendrolycopodium obscurum (L.) A.Haines Ground pine Princess-pine Eastern tree clubmoss Rare clubmoss Flat-branch tree clubmoss Lycopode obscur Native Secure 5 FACU 7048 5 Dendrolycopodium obscurum (L.) A. Haines - flat-branched tree clubmoss - A wide variety of mesic forests and forest opening including forested roadsides. Most common in all but the coolest parts of New York. Appears to do best in successional forests, areas with thin soils, and sites with recent disturbances. LYOB NN as Lycopodium obscurum L. dendrolycopodium/obscurum 204198-Dendrolycopodium-obscurum 17032 407096 200002756 Lycopodium_obscurum Dendrolycopodium obscurum nymap.svg Lycopodium obscurum (homeredwardprice) 001.jpg Dendrolycopodium dendroideum (Michx.) A.Haines 1803 Lycopodium dendroideum Michx. 1890 Lycopodium obscurum L. dendroideum D.C.Eaton 1916 Lycopodium obscurum L. hybridum Farw. 2003 Dendrolycopodium dendroideum (Michx.) A. Haines Prickly tree clubmoss Northern tree clubmoss Treelike clubmoss Prickle tree club-moss Lycopode dendroïde Native Secure 5 FACU 1881 5 LYDE NN as Lycopodium dendroideum Michx. dendrolycopodium/dendroideum 199979-Dendrolycopodium-dendroideum 17027 465751 233500760 Lycopodium_dendroideum Dendrolycopodium dendroideum nymap.svg Dendrolycopodium dendroideum iNat-100397591.jpg Dendrolycopodium hickeyi (W.H.Wagner, Beitel, & Moran) A.Haines 1977 Lycopodium obscurum L. isophyllum Hickey 1989 Lycopodium hickeyi W.H.Wagner, Beitel & R.C.Moran 2003 Dendrolycopodium hickeyi (W.H.Wagner, Beitel, & Moran) A.Haines Pennsylvania clubmoss Hickey's tree club-moss Lycopode de Hickey Native Secure 5 Perennial Herb 6255 5 LYHI2 NN as Lycopodium hickeyi W.H. Wagner, Beitel & Moran dendrolycopodium/hickeyi 204197-Dendrolycopodium-hickeyi 503611 233500761 Lycopodium_hickeyi Dendrolycopodium hickeyi nymap.svg Dendrolycopodium hickeyi iNat-98729163.jpg Spinulum Lycopodium annotinum (8342272166).jpg Spinulum annotinum Spinulum (ˈspīn-yə-ləm) is also treated as Lycopodium sect. Annotina. Lycopodiales Lycopodiaceae Lycopodioideae Spinulum Spinulum Bristly clubmoss 1108 3 Spinulum annotinum (L.) A.Haines 1753 Lycopodium annotinum L. 1772 Lycopodium dubium Zoëga 2003 Spinulum annotinum (L.) A.Haines Common bristly clubmoss Common stiff clubmoss Interrupted clubmoss Lycopode interrompu Native Secure 5 1876 5 LYAN2 NN as Lycopodium annotinum 6594 Lycopodium annotinum spinulum/annotinum 204168-Spinulum-annotinum 17018 465750 200002748 Spinulum annotinum nymap.svg Lycopode au Creux du Van - img 19577.jpg Spinulum canadense (Nessel) A.Haines 1772 Lycopodium dubium auct. non Zoëga 1834 Lycopodium pungens (Desv. ex Spring) Bach.Pyl. ex Iljin 1842 Lycopodium annotinum L. pungens Desv. 1842 Lycopodium pungens La Pylaie ex Desv. 1843 Lycopodium annotinum L. montanum Tuck. 1939 Lycopodium canadense Romhild ex Nessel 1968 Lycopodium annotinum L. pungens Hultén 1969 Lycopodium annotinum L. dubium (Zoëga) Kallio, Laine & Y.Mäkinen 2003 Spinulum canadense (Romhild ex Nessel) A.Haines Northern bristly clubmoss Northern stiff clubmoss Northern interrupted clubmoss Native Endangered 10 1884 1-2 IPNI 6594 Lycopodium annotinum spinulum/canadense 204199-Spinulum-canadense 50305390 Spinulum canadense nymap.svg Spinulum canadense iNat 227208297.jpg Spinulum annotinum × canadense Spinulum annotinum Spinulum canadense Hybrid bristly clubmoss Native Unranked 7168 U Essex county only Spinulum annotinum × canadense nymap.svg Lycopodium annotinum 1127942872.jpg Diphasiastrum Ground cedar (genus Diphasiastrum) is also treated as Lycopodium sect. Complanata. Lycopodiales Lycopodiaceae Lycopodioideae Diphasiastrum Diphasiastrum Ground-cedar 1085 7 Holub (1975) Diphasiastrum digitatum (Dill. ex A.Braun) Holub 1848 Lycopodium digitatum Dill. ex A.Braun 1901 Lycopodium complanatum (L.) Holub flabelliforme Fernald 1911 Lycopodium flabelliforme (Fernald) Blanch. 1965 Diphasium complanatum (L.) Holub flabelliforme (Fernald) Á.Löve & D.Löve in Univ. Colorado Stud. 17: 4 (1965) 1975 Diphasiastrum digitatum (Dill. ex A.Braun) Holub Southern groundcedar Southern ground-cedar Fan club-moss Southern running-pine Lycopode en éventail Courants verts Lycopode aplati variété en éventail Native Secure S5, G5 4 1880 5 LYDI3 NN as Lycopodium digitatum Dill. ex A. Braun 122100-Diphasiastrum-digitatum 6576 Diphasiastrum/digitatum 17028 456047 233500582 Diphasiastrum_digitatum 26610808 1004922-2 Diphasiastrum digitatum nymap.svg Diphasiastrum digitatum6.jpg Diphasiastrum tristachyum (Pursh) Holub 1814 Lycopodium tristachyum Pursh Fl. Amer. Sept. 2: 653 (1813) 1840 Lycopodium clavatum L. tristachyum (Pursh) Hook. Fl. Bor.-Amer. 2: 267 (1840) 1848 Lycopodium complanatum L. tristachyum (Pursh) Dostál Kvĕtena ČSR: 6 (1948) 1859 Lycopodium complanatum L. chamaecyparissus (A.Braun ex Mutel) Milde in Nov. Actorum Acad. Caes. Leop.-Carol. Nat. Cur. 26: 395 (1858 publ. 1859) 1905 Lycopodium complanatum L. tristachyon Clute Fern Allies: 92 (1905) 1944 Diphasium tristachyum (Pursh) Rothm. Feddes Repert. Spec. Nov. Regni Veg. 54: 65 (1944) 1961 Diphasium chamaecyparissus (A. Braun ex Mutel) Á. Löve & D. Löve in Bot. Not. 114: 34 (1961) 1975 Diphasiastrum tristachyum (Pursh) Holub Preslia 47: 108 (1975) Blue ground-cedar Deep-root clubmoss Native Secure S5, G5 4 1877 5 LYTR NN as Lycopodium tristachyum Pursh 122101-Diphasiastrum-tristachyum 147677-2 Diphasiastrum/tristachyum 17037 233500584 Diphasiastrum_tristachyum 26610809 Diphasiastrum tristachyum nymap.svg Diphasiastrum tristachyum 240813d 2x3.jpg Diphasiastrum complanatum (L.) Holub Preslia 47: 108 (1975) 1753 Lycopodium complanatum L. Sp. Pl.: 1104 (1753) 1805 Lepidotis complanata (L.) P.Beauv. Prodr. Aethéogam.: 108 (1805) 1840 Lycopodium anceps Wallr. Linnaea 14: 676 (1840), nom. illeg. 1944 Diphasium complanatum (L.) Rothm. Feddes Repert. Spec. Nov. Regni Veg. 54: 64 (1944) 1958 Diphasium anceps (Asch.) Á.Löve & D.Löve Nucleus (Calcutta) 1(1): 7 (1958) 1975 Diphasiastrum complanatum (L.) Holub Preslia 47: 108 (1975) Northern groundcedar Trailing clubmoss Northern running-pine Lycopode aplati Native Endangered S2, G5 8 1882 1 LYCO3 NN as Lycopodium complanatum L. 82391-Diphasiastrum-complanatum Diphasiastrum/complanatum 512326 456044 200002739 Diphasiastrum_complanatum Diphasiastrum complanatum nymap.svg Diphasiastrum complanatum Острівський зак-к.jpg Diphasiastrum sitchense (Rupr.) Holub (1975) 1845 Lycopodium sitchense Rupr. Beitr. Pflanzenk. Russ. Reiches 3: 30 (1845) 1916 Lycopodium complanatum L. sitchense (Rupr.) Farw. in Rep. (Annual) Michigan Acad. Sci. 18: 94 (1916) 1923 Lycopodium × sabinifolium Willd. sitchense (Rupr.) Fernald in Rhodora 25: 166 (1923) 1965 Lycopodium × sabinifolium Willd. sitchense (Rupr.) Calder & Roy L.Taylor in Canad. J. Bot. 43: 1387 (1965) 1965 Diphasium sitchense (Rupr.) Á.Löve & D.Löve Univ. Colorado Stud., Ser. Biol. 17: 3 (1965) 1975 Diphasiastrum sitchense (Rupr.) Holub Preslia 47: 108 (1975) Sitka groundcedar Alaskan clubmoss Sitka club-moss Lycopode de Sitka Native Endangered S1, G5 10 1878 1 LYSI NN as Lycopodium sitchense Rupr. 204303-Diphasiastrum-sitchense Diphasiastrum/sitchense 512329 465748 233500583 Diphasiastrum sitchense nymap.svg Diphasiastrum sitchense iNat-42883913.jpg Diphasiastrum × sabinifolium (Willd.) Holub (1975) Diphasiastrum sitchense Diphasiastrum tristachyum Lycopodium × sabinifolium Willd. Lycopodium sabinifolium Willd. Lycopodium armatum Desv. Ground-fir Savinleaf groundpine Savin-leaved groundpine Cypress clubmoss Native Unranked SNA, GNA 1875 3 LYSA NN 326170-Diphasiastrum---sabinifolium 819485 456050 233500584 Diphasiastrum × sabinifolium nymap.svg Diphasiastrum × sabinifolium iNat-47838781.jpg Diphasiastrum × zeilleri (Rouy) Holub (1975) Diphasiastrum complanatum Diphasiastrum tristachyum Lycopodium × zeilleri Diphasium zeilleri (Rouy) Damboldt Zeiller's running-pine Zeiller's clubmoss Native Unranked SNA, GNA 7047 2 LYZE NN 326167-Diphasiastrum---zeilleri 503612 233500584 Diphasiastrum × zeilleri nymap.svg Diphasiastrum complanatum (3121381112) 2x3.jpg Diphasiastrum × habereri (House) Holub (1975) Diphasiastrum digitatum Diphasiastrum tristachyum Lycopodium × habereri House Lycopodium complanatum habereri Lycopodium tristachyum habereri Haberer's clubmoss Native Unranked SNA, GNA 6251 U LYHA2 NN as Lycopodium × habereri House 326169-Diphasiastrum---habereri 17029 233500584 Diphasiastrum × habereri nymap.svg Diphasiastrum digitatum (homeredwardprice) 003.jpg Subfamily Lycopodielloideae Lycopodiella Bog clubmoss. (Lycopodiella in Latin means little wolf's-foot.) Lycopodiales Lycopodiaceae Lycopodielloideae Lycopodiella Lycopodiella Bog clubmoss 736 6 Holub (1964) Lycopodiella inundata (L.) Holub 1753 Lycopodium inundatum L. 1805 Plananthus inundatus (L.) P.Beauv. 1852 Lepidotis inundata (L.) Opiz 1912 Lepidotis inundata (L.) Börner 1964 Lycopodiella inundata (L.) Holub Northern bog clubmoss Marsh clubmoss Inundated clubmoss Lycopodielle inondé Native Secure 8 OBL Perennial Subshrub 1874 5 LYIN2 NN lycopodiella/inundata 503609 400775 200002745 26604915 Lycopodiella inundata nymap.svg Lycopodiella inundata 003.jpg Lycopodiella appressa (Chapm.) Cranfill 1843 Lycopodium inundatum L. bigelovii Tuck. 1878 Lycopodium inundatum L. appressum Chapm. 1900 Lycopodium appressum F.E.Lloyd & Underw. 1981 Lycopodiella appressa (Chapm.) Cranfill 1985 Lycopodiella bigelowii (Tuck.) Holub Southern bog clubmoss Appressed bog-clubmoss Chapman's club-moss Native Likely secure 7 FACW Perennial Subshrub 1871 4 LYAP4 NN lycopodiella/appressa 503604 233500756 Lycopodiella appressa nymap.svg BB-0104 Lycopodium adpressum.png Lycopodiella alopecuroides (L.) Cranfill 1753 Lycopodium alopecuroides L. 1944 Lepidotis alopecuroides (L.) Rothm. 1981 Lycopodiella alopecuroides (L.) Cranfill Foxtail bog-clubmoss Foxtail clubmoss Native Vulnerable S2?, G2 7 FACW Perennial Subshrub 1885 3 LYAL5 N0 lycopodiella/alopecuroides 503603 233500755 Lycopodiella alopecuroides nymap.svg Kings, Nassau, Richmond, Suffolk Lycopodiella alopecuroides.jpg Lycopodiella × copelandii (Eiger) Cranfill (1981) L. alopecuroides L. appressa Copeland's bog clubmoss Copeland's clubmoss Native Unranked 7089 U LYCO8 N0 503608 nymap.svg Suffolk Ny hybrid.svg Lycopodiella × gilmanii A.Haines (2003) L. appressa L. inundata Gilman's bog clubmoss Gilman's clubmoss Native Unranked 1872 U IPNI 819486 nymap.svg Ny hybrid.svg Lycopodiella Bog clubmoss 736 X (excluded taxa) Lycopodiella × robusta ( R.J.Eaton) A.Haines (2002) L. alopecuroides L. inundata Lycopodium inundatum var. robustum Bog clubmoss Native Excluded 1873 U IPNI Excluded nymap.svg Ny hybrid.svg Pseudolycopodiella Lycopodiales Lycopodiaceae Lycopodielloideae Pseudolycopodiella Pseudolycopodiella (L.) Holub False bog-clubmoss 1104 1 Pseudolycopodiella caroliniana (L.) Holub 1753 Lycopodium carolinianum L. 1968 Lycopodiella caroliniana (L.) Pic.Serm. 1983 Pseudolycopodiella caroliniana Holub 1983 Pseudolycopodiella meridionalis Holub Carolina clubmoss Slender clubmoss Slender bog club-moss Native Endangered 9 1carolina-clubmoss Pseudolycopodiella caroliniana Endangered S1 G5T4 1889 1 LYCAC N0 not listed in Canada pseudolycopodiella/caroliniana 503606 412269 233500995 26610810 Pseudolycopodiella caroliniana nymap.svg Lycopodiella caroliniana (L.) Pic.Serm. (7858727400).jpg Order Selaginellales Family Selaginellaceae Selaginella Three spikemoss (Selaginella) species have been found growing wild in New York. Selaginellales Selaginellaceae Selaginella Selaginella Spikemoss 149 3 Selaginella rupestris (L.) Spring Flora 21: 182 (1838) 1753 Lycopodium rupestre L. Pl.: 1101 (1753) 1803 Stachygynandrum rupestre (L.) P.Beauv. Prodr. Aethéogam.: 113 (1803) 1838 Selaginella rupestris (L.) Spring Flora 21: 182 (1838) 1849 Selaginella rupestris (L.) Spring borealis Spring Mém. Acad. Roy. Sci. Belgique 24: 57 (1849) 1891 Lycopodioides rupestris (L.) Kuntze Gen. Pl. 2: 827 (1891) 1900 Selaginella bourgeaui Hieron. Hedwigia 39: 295 (1900) 1908 Selaginella rupestris (L.) Spring borealis (Spring) Clute Fern Bull. 16: 52 (1908) 1992 Bryodesma rupestre (L.) Soják Preslia 64: 155 (1992) Rock spikemoss Dwarf spikemoss Northern selaginella Sélaginelle des rochers Sélaginelle rupestre Native Likely secure 9 UPL Perennial Herb S4 6258 4 Dry rock outcrops, ledges, and thin soil over bedrock. SERU NN 133266-Selaginella-rupestris 30039470-2 selaginella/rupestris 17091 200002816 Selaginella rupestris nymap.svg Selaginella rupestris iN-34325628.jpg Selaginella apoda (L.) Spring 1753 Lycopodium apodum L. 1840 Selaginella apus Spring 1840 Selaginella apoda (L.) Spring Meadow spikemoss Creeping spikemoss Sélaginelle apode Native Unranked Under review2019 Rare Plant Status Lists – Review List - New York Natural Heritage Program 6 FACW 6257 2? SEAP N0 Not found in Canada selaginella/apoda 130887-Selaginella-apoda 17066 200002770 Selaginella apoda nymap.svg Selaginella apoda iN-11291436.jpg Selaginella eclipes W.R.Buck 1977 Selaginella eclipes W.R.Buck 1996 Selaginella apoda eclipes Skoda Hidden spikemoss Buck's meadow spikemoss Creeping spikemoss Native Endangered 8 FACW 7125 1 No specimens SEEC NN selaginella/eclipes 17079 233501230 26602874 Selaginella eclipes nymap.svg Selaginella eclipes iNat-19576466.jpg Selaginella Spikemoss 149 X (excluded taxa) Selaginella selaginoides (L.) P. Beauv. ex Mart. & Schrank 1753 Lycopodium selaginoides L. 1805 S. spinosa P.Beauv. illeg. 1841 S. selaginoides (L.) P.Beauv. ex Mart. & Schrank Club spikemoss USDA NRCS National Plant Data Team Northern spikemoss Flora Novae Angliae Northern spike-moss ARS-GRIN: FNA Editorial Committee 1994 Prickly mountain-moss ARS-GRIN: FNA Editorial Committee 1994 Lesser-club-moss ARS-GRIN: Botanical Society of the British Isles 2007 N. America native Excluded FACW Perennial Herb X 635 EXCLUDED SESE NN selaginella/selaginoides 412259 Excluded nymap.svg Selaginella selaginoides02.jpg Order Isoetales Family Isoetaceae Isoetes Isoetes echinospora 74924873 (4x3).jpg Isoetes echinospora North American Isoetes lacustris plants have recently been segregated as Isoetes macrospora but can't be easily differentiated except by geography.W. Carl Taylor, Neil T. Luebke, Donald M. Britton, R. James Hickey, Daniel F. Brunton (2020). FNA Volume 2. Flora of North America Association Isoetales Isoetaceae Isoetes Isoetes L. Quillwort 241 9 Isoetes engelmannii A.Braun 1846 Isoetes engelmannii A.Braun Flora 29: 178 (1846) 1846 Isoetes lacustris L. microspora Engelm. A. Braun, Flora 29: 179 (1846) 1867 Isoetes engelmannii A.Braun gracilis Engelm. A.Gray, Manual, ed. 5: 677 (1867) 1891 Calamaria engelmannii (A.Braun) Kuntze Revis. Gen. Pl. 2: 828 (1891) 1905 Isoetes engelmannii A.Braun fontana A.A.Eaton Fern Bull. 13: 52 (1905) 1994 Isoetes × fairbrothersii J.D.Montgom. & W.C.Taylor Amer. Fern J. 84(4): 120 (1994) Engelmann's quillwort Appalachian quillwort Isoète d'Engelmann Native Secure 6 OBL Perennial Herb, Subshrub diploid (2n = 22) S5 7051 5 ISEN NN B9QTW 2688326 164160-Isoetes-engelmannii 30212032-2 0001108952 41 6216 isoetes/engelmannii 17127 250076852 Isoëtes_engelmannii 26602042 30212032-2 Isoetes+engelmannii Isoetes%20engelmannii Isoetes engelmannii nymap.svg 23 counties Cortland (2019) Isoetes engelmannii NRCS-3x4.jpg Isoetes echinospora Durieu muricata A.Löve & D.LöveThe New York Flora Atlas and New England Wild Flower Society list the spiny-spore quillwort as Isoëtes echinospora ssp. muricata because the leaves of the North American plants of bear stomata, while those of the European plants do not. The other sources used here treat ssp. muricata as a synonym. 1789 Isoetes setacea Lam. p.p. 1827 Isoetes tenella Léman ex Desv. POWO Mém. Soc. Linn. Paris 6: 179 (1827) 1864 Isoetes braunii Durieu Bull. Soc. Bot. France 11: 101 (1864) 1867 Isoetes echinospora Durieu braunii Engelm. 1945 Isoetes muricata Durieu braunii (Engelm.) C.F.Reed Amer. Fern J. 35: 83 (1945) 1961 Isoetes echinospora Durieu muricata (Durieu) Á.Löve & D.Löve NYFA Spiny-spore quillwort Spring quillwort Braun's quillwort Bristlelike quillwort Isoète à spores épineuses Isoète à spores spinuleuses Native Likely secure 7 OBL Perennial Herb diploid (2n = 22) S4 7049 4 ISTE5 ? as Isoetes tenella Léman 6215 isoetes/echinospora 17124 412272 250076846 26600171 Isoetes echinospora nymap.svg Isoetes echinospora iNat-25264828q.jpg Isoetes tuckermanii A.Braun ex Engelm. 1867 I. tuckermanii A.Braun ex Engelm. 1900 I. tuckermanii borealis A.A.Eaton 1981 I. acadiensis Kott 1991 I. tuckermanii acadiensis Taylor & Luebke Tuckerman's quillwort Isoète de Tuckerman Native Likely secure 10 tetraploid (2n = 44) 7057 4 ISTU NN 68 isoetes/tuckermanii 17142 250076853 Isoetes tuckermanii nymap.svg Isoetes tuckermanii 15-p.bot-isoe.tucke-007.jpg Isoetes × eatonii R.Dodge Isoetes echinospora Isoetes engelmannii 1896 Isoetes eatonii R.Dodge 1900 Isoetes gravesii A.A.Eaton Isoetes eatonii gravesii A.A.Eaton Eaton's quillwort Native Vulnerable 7050 3 ISEA NN CT,MA,NH,NJ,NY,PA,VT,ON (only) 17123 Isoetes × eatonii nymap.svg Isoetes × eatonii BB-1913-1.png Isoetes macrospora Durieu 1753 Isoetes lacustris L. Sp. Pl.: 1100 (1753) 1864 Isoetes macrospora Durieu Bull. Soc. Bot. France 11: 101 (1864) 1900 Isoetes hieroglyphica A.A.Eaton 1922 Isoetes macrospora L. hieroglyphica N.Pfeiff. Ann. Missouri Bot. Gard. 9: 194 (1922) Big-spore quillwort 2024 iNat Lake quillwort Carved quillwort Warty Quillwort Isoète lacustre Isoète hiéroglyphique Native Rare 10 OBL Perennial Herb decaploid (2n = 110) S3 7054 3 ISLA NN 2688270 164168-Isoetes-macrospora isoetes/lacustris 17131 250076848 Isoëtes_lacustris Isoetes lacustris nymap.svg 5 counties St. Lawrence only Isoetes lacustris rivnenskyi.jpg Isoetes septentrionalis D.F.Brunt. Daniel F. Brunton, in D.F. Brunton & J.M. McNeill (2015), Status, distribution, and nomenclature of Northern Quillwort, Isoetes septentrionalis (Isoetaceae) in Canada. The Canadian Field-Naturalist. Vol 129, No 2 (2015) 1846 Isoetes riparia Engelm. ex A.Braun canadensis Engelm. Trans. Acad. Sci. St. Louis 4: 383 (1882) misapplied 1903 Isoetes canadensis robbinsii A.A. Eaton 2015 Isoetes septentrionalis D.F. Brunt. Canad. Field-Naturalist 129: 176 (2015) Northern shore quillwort Northern quillwort isoète du Nord Native Endangered 10 OBL Perennial Herb tetraploid (2n = 44) 7055 1 Isoetes septentrionalis D.F. Brunt. - northern shore quillwort ISRIA N0 as Isoetes riparia var. amesii 7055 29176 isoetes/riparia 17140 250076855 Isoetes riparia var canadensis nymap.svg BB-0123 Isoetes riparia.png Isoetes × robusta (Engelm.) D.F.Brunt. Isoetes engelmannii Isoetes septentrionalis 1898 Isoetes × dodgei A.A.Eaton auct Isoetes × brittonii auct Isoetes engelmannii × riparia Engelmann's quillwort northern shore quillwort Native No recent reports 7052 Z? Ulster County only (1990 list) ISBR5 N0 CT, NJ, NY, PA, SC, VA (only) isoetes/ 503213 412271 Isoetes × brittonii nymap.svg BB-0127 Isoetes engelmanni.png Isoetes (excluded) Isoetales Isoetaceae Isoetes Isoetes Quillwort 241 8 (excluded taxa) Isoetes riparia Engelm. ex A.BraunIsoetes riparia was apparently misapplied to reports of Isoetes septentrionalis in New York. (NYFA (2016) 1846 Isoetes riparia Engelm. ex A.Braun 18?? Isoetes saccharata Engelm. in A.Gray 1891 Calamaria saccharata (Engelm.) Kuntze 1901 Isoetes canadensis (Engelm.) A.A.Eaton 1949 Isoetes riparia saccharata ( Engelm. ex A.Gray ) Proctor Shore quillwort Canada shore quillwort Native Excluded X 131 ISRI NN Isoetes/riparia 17140 26602056 Isoetes%20riparia Excluded nymap.svg BB-0123 Isoetes riparia.png Isoetes × harveyi A.A.Eaton Isoetes lacustris Isoetes tuckermanii 1900 Isoetes × heterospora A.A.Eaton Isoetes tuckermanii harveyi Clute Harvey's quillwort Native Excluded X 631 ISHA4 NN isoetes/tuckermanii 536383 250076853 Excluded nymap.svg Ny hybrid.svg Isoetes × novae-angliae D.F. Brunt. & D.M. Britton Isoetes septentrionalis Isoetes tuckermanii New England quillwort Native Excluded X 632 ISNO N0 isoetes/tuckermanii 536383 250076853 Excluded nymap.svg Ny hybrid.svg