Sensory Systems/Auditory System

3 Introduction The sensory system for the sense of hearing is the auditory system. This wikibook covers the physiology of the auditory system, and its application to the most successful neurosensory prosthesis - cochlear implants. The physics and engineering of acoustics are covered in a separate wikibook, Acoustics. An excellent source of images and animations is "Journey into the world of hearing" Journey into the world of hearing http://www.cochlea.org/en/spe NeurOreille and authors 2010 . The ability to hear is not found as widely in the animal kingdom as other senses like touch, taste and smell. It is restricted mainly to vertebrates and insects. Within these, mammals and birds have the most highly developed sense of hearing. The table below shows frequency ranges of humans and some selected animals: Humans 20-20'000 Hz Whales 20-100'000 Hz Bats 1'500-100'000 Hz Fish 20-3'000 Hz The organ that detects sound is the ear. It acts as receiver in the process of collecting acoustic information and passing it through the nervous system into the brain. The ear includes structures for both the sense of hearing and the sense of balance. It does not only play an important role as part of the auditory system in order to receive sound but also in the sense of balance and body position. thumb|Mother and child thumb|Humpback whales in the singing position thumb|Big eared townsend bat thumb|Hyphessobrycon pulchripinnis fish Humans have a pair of ears placed symmetrically on both sides of the head which makes it possible to localize sound sources. The brain extracts and processes different forms of data in order to localize sound, such as: the shape of the sound spectrum at the tympanic membrane (eardrum) the difference in sound intensity between the left and the right ear the difference in time-of-arrival between the left and the right ear the difference in time-of-arrival between reflections of the ear itself (this means in other words: the shape of the pinna (pattern of folds and ridges) captures sound-waves in a way that helps localizing the sound source, especially on the vertical axis. Healthy, young humans are able to hear sounds over a frequency range from 20 Hz to 20 kHz.November 2022 We are most sensitive to frequencies between 2000 and 4000 HzNovember 2022 which is the frequency range of spoken words. The frequency resolution is 0.2% which means that one can distinguish between a tone of 1000 Hz and 1002 Hz. A sound at 1 kHz can be detected if it deflects the tympanic membrane (eardrum) by less than 1 AngstromNovember 2022, which is less than the diameter of a hydrogen atom. This extreme sensitivity of the ear may explain why it contains the smallest bone that exists inside a human body: the stapes (stirrup). It is 0.25 to 0.33 cm long and weighs between 1.9 and 4.3 mg.November 2022 The following video provides an overview of the concepts that will be presented in more detail in the next sections. 550px|center|thumb|This animated video illustrates how sounds travel to the inner ear, and then to the brain, where they are interpreted and understood. The cochlea in the inner ear is a spiral-shaped organ that contains hair cells, which sense sound vibrations. Hair cells convert sound vibrations into chemical signals that the auditory nerve can understand. Visual System Vestibular System Sensory_Systems