Flora of New York/Rosoideae 1

Rosales : Rosaceae : Rosoideae 1 : Ulmarieae, Rubeae, Colurieae, Sanguisorbeae Fabales Rosoideae 2 Rosales: Rosaceae: Rosoideae 2 Rosales introduction The order Rosales is part of the nitrogen-fixing clade in the fabids: fabids Zygophyllales COM clade Celastrales Malpighiales Oxalidales nitrogen‑fixing clade  Fabales Rosales Cucurbitales Fagales Rosaceae introduction Rosoideae introduction The Rosaceae subfamily Rosoideae contains the following tribes: Agrimonieae (Acaena, Agrimonia, Poteridium, Poterium, Sanguisorba) Colurieae (Fallugia, Geum, Sieversia, Waldsteinia) Potentilleae (Alchemilla, Aphanes, Chamaerhodos, Comarum, Dasiphora, Drymocallis, Duchesnea, Fragaria, Horkelia, Horkeliella, Ivesia, Potentilla, Sibbaldia, Sibbaldiopsis) Roseae (Rosa) Rubeae (Rubus) Ulmarieae (Filipendula) Tribe Ulmarieae Filipendula Filipendula rubra 2017-06-25 3209.jpg Filipendula rubra Filipendula contains about ten Eurasian species and two North American species. Of these, one Midwestern and two Eurasian species have been found to have naturalized in New York. Filipendula rubra (queen of the prairie) is the only North American native Filipendula found in New York State, where it is only known to be present in the wild as a garden escape.Ivan A. Schanzer (2020). Flora of North America, Volume 9. Page 25. Flora of North America Association. 5 Nov. 2020 It can be found on roadsides, ditches, thickets, and disturbed soils, but it has been less common than the Eurasian Filipendula ulmaria.Werier, David, et al. (2023). Filipendula rubra in New York Flora Atlas. New York Flora Association, Albany, New York. 12 Dec. 2023. Rosales Rosaceae Rosoideae Ulmarieae Filipendula Filipendula Queen, Dropwort 948 3 Filipendula rubra (Hill) B.L.Rob. Benjamin Lincoln Robinson (1864-1935). Rhodora 8(94):204. 1906. 1768 Ulmaria rubra Hill 1770 Spiraea lobata Gronov. ex Jacq. 1906 Filipendula rubra B.L.Rob. 1836 Thecanisia angustifolia Raf. 1836 Thecanisia purpurea Raf. 1837 Thecanisia lobata (Gronov.) Raf. 1879 Filipendula lobata (Gronov.) Maxim. 1891 Spiraea rubra (Hill) Britton 1924 Filipendula lobata (Gronov.) Maxim. albiflora Bergmans 1924 Filipendula lobata (Gronov.) Maxim. venusta-magnifica Bergmans Queen-of-the-prairie Prairie dropwort Red queen-of-the-meadow Filipendule rouge Introduced US midwest N. America native Cultivated in NY 0 FACW Perennial Herb-forb Part shade G4-G5 2695 X Roadsides, thickets, ditches, and disturbed soils. Much less common than the closely related F. ulmaria. FIRU2 NX 104799-2 8748 filipendula/rubra 25170 456822 Filipendula_rubra 27800864 FIRU2 Filipendula+rubra Filipendula%20rubra Filipendula rubra nymap.svg 13 counties 25 counties Filipendula rubra inflorescence.JPG 120 Filipendula ulmaria (L.) Maxim. Trudy Imp. S.-Peterburgsk. Bot. Sada 6: 251 (1879) 1753 Spiraea ulmaria L. 1819 Spiraea denudata J.& C.Presl 1822 Spiraea ulmaria L. tomentosa Hayne) 1838 Thecanisia ulmaria (L.) Raf. 1879 Filipendula ulmaria Maxim. 1879 Filipendula ulmaria Maxim. denudata (J.Presl & C.Presl) Maxim. 1889 Filipendula denudata Fritsch 1889 Spiraea ulmaria L. tomentosa (Hayne) St.-Lag. (not validly publ.) 1894 Ulmaria ulmaria (L.) Barnhart (not validly publ.) 1895 Thecanisia ulmaria (L.) Raf. ex B.D.Jacks. (comb. not validly publ.) 1909 Filipendula ulmaria Maxim. denudata Hayek Queen-of-the-meadow Meadowsweet Meadow dropwort Elm queen-of-the-meadow White queen-of-the-meadow Reine-des-prés Introduced temperate Eurasia Potentially invasive Note:Current invasive species rule revisions propose to regulate Filipendula ulmaria as Restricted in Wisconsin.Pfingsten, I.A., D. Rose, 2023, Filipendula ulmaria (L.) Maxim.: U.S. Geological Survey, Nonindigenous Aquatic Species Database, Gainesville, FL, https://nas.er.usgs.gov/queries/FactSheet.aspx?SpeciesID=3810, Revised 8/30/2023, Accessed 12/16/2023 14017 4 FAC FACW Perennial Herb-forb 2694 X FIUL XX 2987999 63134-Filipendula-ulmaria 724943-1 8749 filipendula/ulmaria 25171 17105 27800269 Filipendula ulmaria nymap.svg 21 counties 16 counties 20150811Filipendula ulmaria2.jpg Filipendula vulgaris Moench 1753 Spiraea filipendula L. 1768 Ulmaria filipendula (L.) Hill 1782 Filipendula hexapetala Gilib. (opus utique oppr.) 1794 Filipendula vulgaris Moench 1822 Spiraea vulgaris (Moench) Gray (nom. superfl.) 1824 Spiraea filipendula L. vulgaris Cambess. 1879 Filipendula hexapetala Gilib. superfl. 1788 Ulmaria filipendula (L.) Hill ex Focke 1894 Filipendula filipendula (L.) Voss (not validly publ.) 1915 Ulmaria angustifolia Rehder Dropwort Common queen-of-the-meadow Filipendule vulgaire Filipendule Introduced temperate Eurasia Not naturalized 2708 X FIVU XX 2988069 62020-Filipendula-vulgaris 724945-1 8750 filipendula/vulgaris 25172 104611 Filipendula_vulgaris 27800060 Filipendula vulgaris nymap.svg Madison, Chemung (1938) No RG observations Rosaceae - Filipendula vulgaris-1 (8303629717).jpg Tribe Rubeae The Rubeae include the genera Rubus and Dalibarda (which is here treated as a monospecific subgenus of Rubus. Rubus Rubus subg. Chamaemorus Rosaceae- Rubus chamaemorus (Cloudberry) 2.jpg Rubus chamaemorus The native status of cloudberry (Rubus chamaemorus) in New York could be subject to question. It is a circumboreal bramble, native to all provinces of Canada and present in the states of Alaska, Minnesota, New Hampshire, and Maine. However, the only vouchered report of cloudberry growing outside of cultivation in New York was from Montauk in Suffolk County, Long Island in 1908.Eric E. Lamont and Stephen M. Young (2005) "Juncus diffusissimus, an addition to the flora of New York, with notes on its recent spread in the United States." The Journal of the Torrey Botanical Society, 132(4): 635-643. 2005.H. K. Svenson (1936) "The Early Vegetation of Long Island," Brooklyn Botanical Garden Record 25: 207-227. Republished in the Long Island Botanical Society Newsletter, p.2, 1994. Rosales Rosaceae Rosoideae Rubeae Rubus Chamaemorus Rubus Cloudberry 843 30 Chamaemorus Rubus chamaemorus L. 1753 Rubus chamaemorus L. 1906 Chamaemorus anglica Greene 1906 Chamaemorus norwegica Greene 1920 Chamaemorus chamaemorus (L.) House (not validly publ.) Cloudberry Baked-apple-berry Salmon-berry Malka Chicouté Ronce petit-mûrier Mûres blanches Native No recent reports Impersistent 10 FACW Perennial Herb-subshrub Sun 6526 Z Suffolk county only RUCH NN 8928 rubus/chamaemorus 24850 32286 27800175 RUCH Rubus chamaemorus nymap.svg Suffolk (1908) No NY observations Hjortron.png Rubus subg. Dalibarda Rubus repens 15-p.bot-dali.repens-6.jpg Rubus repens Dalibarda has been treated as a monospecific genus, but the Flora of North America Editorial Committee (2014)Flora of North America Editorial Committee. 2014. Magnoliophyta: Picramniaceae to Rosaceae. Fl. N. Amer. 9: i–xxiv, 1–713. Listed in Tropicos.org. Missouri Botanical Garden. 05 Jul 2015 (http://www.tropicos.org/Name/50334377) treats it as a subgenus of Rubus. Its leaves superficially resemble those of violets, but its flowers and fruit are clearly rubusesque. There continues to be disagreement about whether R. dalibarda or R. repens should be the accepted species name. Rubus repens was found to be invalid but has been proposed for conservation as the accepted name.Turner, I.M. (2023), (2980) Proposal to conserve the name Rubus repens (L.) Kuntze against R. repens Weston (Rosaceae). TAXON, 72: 942-942. https://doi.org/10.1002/tax.12999 Subgenus Dalibarda does appear to form a strongly-supported clade with the rest of Rubus and may nest within Rubus.Alice, L.A. and Campbell, C.S. (1999), Phylogeny of Rubus (rosaceae) based on nuclear ribosomal DNA internal transcribed spacer region sequences . Am. J. Bot., 86: 81-97. https://doi.org/10.2307/2656957 Rosales Rosaceae Rosoideae Rubeae Rubus Dalibarda Rubus Dewdrop 685 1 Dalibarda Rubus repens (L.) Kuntze Revis. Gen. Pl. 1: 223. 1891. 1753 Dalibarda repens L. 1762 Rubus dalibarda L. (nom. superfl.) 1770 Rubus repens Weston (nom. rej. prop.) 1803 Dalibarda violioides Michx. 1806 Dalibarda cordata Stephan (nom. superfl.) 1879 Rubus dalibarda L. caespitosus Kuntze 1879 Rubus dalibarda L. laciniostipulatus Kuntze 1891 Rubus repens (L.) Kuntze (nom. cons. prop.) Dewdrop Robin runaway False-violet Star-violet Dalibarde rampante Native Likely secure 9 FAC Perennial Herb-forb Part shade - shade 6508 4 Rubus repens (L.) Kuntze - dewdrop - Cool hardwood and coniferous forests as well as hummocks in forested swamps. Predominately in shaded habitats with wet-mesic soils. DARE NN Dalibarda repens L. 4TJW9 2999946 223505-2 0000988537 8727 rubus/dalibarda 924430-Rubus-repens 520495 470540 250100445 Rubus_repens 50334377 DARE Rubus+repens Rubus%20dalibarda nymap.svg Dalibarda repens 4284 (2x3).JPG Rubus subg. Anoplobatus Rubus odoratus G4.JPG Rubus odoratuspurple-flowering raspberry The thornless Anoplobatus subgenus contains Rubus odoratus. Rosales Rosaceae Rosoideae Rubeae Rubus Anoplobatus Rubus Raspberry 843 30 Anoplobatus Rubus odoratus L. 1753 Rubus odoratus L. 1892 Rubus odoratus L. columbianus Millsp. 1903 Rubacer odoratum Rydb. 1903 Rubacer columbianum Rydb. 1906 Bossekia odorata Greene 1912 Bossekia columbiana A.Heller Purple-flowering raspberry Purple-flowered raspberry Virginia raspberry Thimbleberry Ronce odorante Framboisier sauvage Calottes Native Secure 7 unranked Perennial Subshrub Sun - shade 2759 5 Forest edges, talus slopes, mesic rocky outcrops, disturbed soils in forests, and thickets. Generally in wet-mesic or sometimes mesic soils and often in at least partly shaded habitats. RUOD NN 8941 rubus/odoratus 24997 32402 250100440 27801320 RUOD Rubus+odoratus Rubus%20odoratus Rubus-odoratus-flower.JPG Rubus subg. Idaeobatus Fertődi zenit málna.jpg Rubus idaeusred raspberry The Rubus subgenus Idaeobatus species are generally referred to as raspberries. Rosales Rosaceae Rosoideae Rubeae Rubus Idaeobatus Rubus Raspberry 843 30 Idaeobatus Rubus idaeus L. strigosus (Michx.) Focke 1803 Rubus strigosus Michx. 1872 Rubus idaeus L. strigosus (Michx.) Maxim. 1906 Batidaea acalyphacea Greene 1906 Batidaea strigosa (Michx.) Greene 1907 Rubus egglestonii Blanch. 1911 Rubus idaeus L. strigosus (Michx.) Focke 1912 Batidaea egglestonii (Blanch.) A.Heller 1913 Rubus acalyphaceus (Greene) Rydb. 1913 Rubus carolinianus Rydb. 1913 Rubus arizonicus (Greene) Rydb. 1913 Rubus subarcticus (Greene) Rydb. 1945 Rubus aliceae L.H.Bailey American red raspberry North American red raspberry Grayleaf red raspberry Framboisier sauvage Native Secure 3 FACU FAC Perennial Shrub S5, G5-T5 2755 5 as Rubus idaeus ssp. strigosus RUIDS2 NN as Rubus idaeus ssp. strigosus BNZCD 2993119 223189-2 0001006872 8934 rubus/idaeus 524636 467362 nymap.svg Texter Mountain Nature Preserve (Revisited) (13) (9387620458).jpg Rubus idaeus L. idaeus 1753 Rubus idaeus L. 1770 Rubus cannabifolius Weston 1770 Rubus elegantissimus Weston 1770 Rubus idaeus L. albus Weston 1770 Rubus idaeus L. biflorus Weston 1770 Rubus idaeus L. laevis Weston 1789 Rubus daeus ruber Aiton 1796 Rubus fragrans Salisb. 1811 Rubus obtusifolius Willd. 1915 Batidaea idaea (L.) Nieuwl. 1915 Batidaea vulgaris Nieuwl. European red raspberry Framboisier Introduced Eurasia FACU FAC Perennial Shrub SNA, G5-T5 2746 X RUIDI XX 77232556-1 0001006864 rubus/idaeus 524635 32348 nymap.svg Fertődi zamatos málna 2.jpg Rubus occidentalis L. Sp. Pl. 493 (1753) 1753 Rubus occidentalis L. 1794 Rubus tomentosus Borkh. (nom. superfl.) 1828 Rubus idaeus L. americanus Torr. 1839 Rubus occidentalis L. incisus Schltdl. 1874 Rubus occidentalis L. euoccidentalis Focke 1890 Rubus occidentalis L. japonicus Miyabe 1891 Rubus occidentalis L. grandiflorus Focke 1898 Rubus fruticosus L. tomentosus Fiori 1902 Rubus occidentalis L. pallidus L.H.Bailey 1905 Rubus occidentalis L. flavobaccus Blanch. 1906 Melanobatus occidentalis (L.) Greene Black raspberry Black cap Thimbleberry Framboisier de Virginie Native Secure 3 Perennial Shrub S5, G5 2751 5 RUOC NN 4TL3H 223350-2 8940 rubus/occidentalis 24854 32401 Rubus%20occidentalis 2018-06-15 10 27 01 Black raspberries ripening along a walking path in the Franklin Farm section of Oak Hill, Fairfax County, Virginia.jpg Rubus × neglectus Peck (pro sp.) Rubus occidentalis Rubus idaeus ssp. strigosus Purple wild raspberry Native Not assessed Perennial Vine, Shrub SNA, GNA 2702 2? RUIDS2 NN as Rubus idaeus ssp. strigosus 24989 nymap.svg Nidda Fauerbach Rubus idaeus ws.png Rubus phoenicolasius Maxim. 1872 Rubus phoenicolasius Maxim. Wineberry Japanese wineberry Wine raspberry Introduced temperate Asia Highly invasive 86%Rubus phoenicolasius High 86 2720 X RUPH X0 CP-4CP4 NE-4NE4 rubus/phoenicolasius 25017 32416 Rubus phoenicolasius nymap.svg Japanse wijnbes rijpe vruchten.jpg Rubus illecebrosus Focke 1899 Rubus illecebrosus Focke Strawberry raspberry Charming raspberry Framboisier fraisier Introduced eastern Asia Potentially invasive 2785 X RUIL XX IO rubus/illecebrosus 504856 32354 Rubus illecebrosus - fruit (aka).jpg Rubus parvifolius L. 1753 Rubus parvifolius L. 1784 Rubus triphyllus Thunb. 1833 Rubus purpureus Bunge Japanese raspberry 2021 NYFA Small-leaf bramble 2021 iNaturalist Trailing raspberry 1961 Webster's third new international dictionary. Introduced Asia Australia Potentially invasive Naturalized X SNA 7605 Xn 51645-Rubus-parvifolius rubus/parvifolius 32409 27803708 nymap.svg NYFA: 1 county Rubus parvifolius 0907.JPG Rubus subg. Rubus Subgenus Rubus contains the blackberries and dewberries. Rubus subg. Rubus sect. Allegheniensis 2016-07-25 16 21 11 Blackberries along the Blue Ridge Parkway in Deep Gap, Yancey County, North Carolina.jpg Rubus allegheniensis Rubus L. sect. Allegheniensis contains Rubus allegheniensis, R. alumnus, and R. pennus.Rubus L. sect. Allegheniensis in USDA-ARS, GRIN-Global, U.S. National Plant Germplasm System. 23 September 2023 Of these, only R. allegheniensis is listed for New York State. Rosales Rosaceae Rosoideae Rubeae Rubus Rubus Allegheniensis Rubus Blackberry 843 30 Rubus Allegheniensis Rubus allegheniensis Porter 1890 Rubus villosus Aiton not Thunb. montanus Porter 1894 Rubus montanus (Porter) Porter (later homonym) 1896 Rubus allegheniensis Porter 1906 Rubus pergratus Blanch. 1906 Rubus amnicola Blanch. 1945 Rubus avipes L.H.Bailey Allegheny blackberry Common blackberry Highbush blackberry Ronce des Alléghanys Mûrier Ronce alléghanienne Native Secure 3 FACU Perennial Shrub 2747 5 RUAL NN 8920 rubus/allegheniensis 24866 32254 242417156 27801178 Rubus allegheniensis NY-dist-map.png Rubus allegheniensis NRCS-015.jpg Rubus subg. Rubus sect. Canadenses Rubus canadensis (3813766870).jpg Rubus canadensis Rubus L. sect. Canadenses contains Rubus canadensis and R. kennedyanus.Rubus L. sect. Canadenses in USDA-ARS, GRIN-Global, U.S. National Plant Germplasm System. 23 September 2023 Of these, only Rubus canadensis is listed for New York State. Rosales Rosaceae Rosoideae Rubeae Rubus Rubus Canadenses Rubus Blackberry 843 30 Rubus Canadenses Rubus canadensis L. 1753 Rubus canadensis L. 1891 Rubus millspaughii Britton 1895 Selnorition canadensis (L.) Raf. ex B.D.Jacks. 1901 Rubus randii (L.H.Bailey) Rydb. 1931 Rubus kennedyanus Fernald 1932 Rubus montensis L.H.Bailey 1944 Rubus acridens L.H.Bailey 1944 Rubus ulterior L.H.Bailey 1947 Rubus besseyi L.H.Bailey Smooth blackberry Smooth bramble Thornless blackberry Native Vulnerable 2777 3? RUCA16 NN Rubus/canadensis 32281 27801823 Rubus canadensis perrons black.jpg Rubus subg. Rubus sect. Arguti Rubus pensilvanicus flowers.JPG Rubus pensilvanicus The Flora of North America Editorial Committee (2014) treats the Yankee blackberry (R. frondosus), Burnham's blackberry (R. burnhamii), and the prickly Florida blackberry (R. argutus) as being among the 72 synonyms of the Pennsylvania blackberry (R. pensilvanicus).Rubus pensilvanicus Poir. at Tropicos.org. Missouri Botanical Garden. 04 Jul 2015 <http://www.tropicos.org/Name/27801334>Rubus pensilvanicus Poir. at Canadensys, VASCAN last updated on 2015-06-29 ITIS also lists Rydberg's blackberry (R. rydbergianus), along with the other three, as a synonym of (R. pensilvanicus).Rubus pensilvanicus Poir. at Integrated Taxonomic Information System (ITIS) (http://www.itis.gov). 04 Jul 2015 This leaves sect. Arguti equivalent to Rubus pensilvanicus itself. Rosales Rosaceae Rosoideae Rubeae Rubus Rubus Arguti Rubus Blackberry 843 30 Rubus Arguti Rubus pensilvanicus Poir. J.B.A.M.de Lamarck, Encycl. 6: 246 (1804) 1804 Rubus pensilvanicus Poir. 1822 Rubus argutus Link 1824 Rubus frondosus Bigelow 1913 Rubus brainerdii Rydb. 1932 Rubus abactus L.H.Bailey 1933 Rubus penetrans L.H.Bailey 1945 Rubus barbarus L.H.Bailey 1945 Rubus bellobatus L.H.Bailey 1945 Rubus blakei L.H.Bailey 1945 Rubus burnhamii L.H.Bailey 1945 Rubus rydbergianus L.H.Bailey Pennsylvania blackberry Pennsylvania raspberry Ronce de Pennsylvanie Native Secure 3 2749 5 RUPE3 NN 4TM7H 2989061 30263954-2 0001007097 78894-Rubus-pensilvanicus 8944 rubus/pensilvanicus 504874 104942 242417169 Rubus_pensilvanicus 27801334 Rubus pensilvanicus iNat 3654011.jpg Rubus subg. Rubus sect. Cuneifolii Rubus cuneifolius NRCS-1.jpg Rubus cuneifolius Rosales Rosaceae Rosoideae Rubeae Rubus Rubus Cuneifolii Rubus Blackberry 843 30 Rubus Cuneifolii Rubus cuneifolius Pursh Fl. Amer. Sept. 1: 347 (1814) 1788 Rubus parvifolius Walter (sensu auct.) 1814 Rubus cuneifolius Pursh 1925 Rubus cuneifolius Pursh angustior L.H.Bailey 1932 Rubus audax L.H.Bailey 1938 Rubus longii Fernald 1941 Rubus cuneifolius Pursh subellipticus Fernald 1943 Rubus chapmanii L.H.Bailey 1943 Rubus georgiensis L.H.Bailey 1943 Rubus humei L.H.Bailey 1943 Rubus randolphiorum L.H.Bailey 1943 Rubus probativus L.H.Bailey 1970 Rubus dixiensis H.A.Davis, A.M.Fuller & T.Davis Sand blackberry Native Endangered No recent reports 10 Perennial Shrub 2758 1 RUCU N0 Endangered in CT, NH, NY, PA 4TJVP 2996841 128854-Rubus-cuneifolius 30201781-2 0001006778 24905 32304 Rubus cuneifolius NY-dist-map.png 3 counties 2 counties Sand-Blackberry-flower (5622475257) (3x4).gif Rubus subg. Rubus sect. Setosi The Flora of North America Editorial Committee (2014) treats both the Bristly Oswego blackberry (R. dissimilis) and the Vermont blackberry (R. vermontanus) as being among the 40 synonyms of the Bristly blackberry (R. setosus).Rubus setosus Bigelow at Tropicos.org. Missouri Botanical Garden. 04 Jul 2015Rubus setosus Bigelow at Canadensys, VASCAN last updated on 2015-06-29 Rosales Rosaceae Rosoideae Rubeae Rubus Rubus Setosi Rubus Blackberry 843 30 Rubus Setosi Rubus setosus Bigelow Fl. Boston.: 198 (1824) 1824 Rubus setosus Bigelow 1901 Rubus nigricans Rydb. (nom. illeg.) 1904 Rubus vermontanus Blanch. 1904 Rubus vermontanus Blanch. viridifolius Blanch. 1906 Rubus groutianus Blanch. 1907 Rubus semisetosus Blanch. 1932 Rubus wheeleri L.H.Bailey 1941 Rubus dissimilis L.H.Bailey 1941 Rubus lawrencei L.H.Bailey 1947 Rubus beatus L.H.Bailey 1947 Rubus condignus L.H.Bailey 1947 Rubus viridifrons L.H.Bailey Bristly blackberry Setose blackberry Small bristleberry Bristly raspberry Ronce sétuleuse Native Likely secure 2 FACW Perennial Shrub S4 6536 4 RUSE NN 4TM7S 7769228 168165-Rubus-setosus 740337-1 0001007245 8947 rubus/setosus 25046 104974 250100446 27801373 Rubus setosus as Rubus nigricans BB-1913.png Rubus subg. Rubus sect. Rubus Rubus laciniatus 06.jpg Rubus laciniatus Sect. Rubus contains somewhat invasive Old World species of Rubus subg. Rubus. The primary species of this section in New York State is Rubus laciniatus (cutleaf blackberry). The rare-in-NY but possibly naturalizing and invasive Himalayan blackberry (Rubus bifrons and/or Rubus armeniacus), listed in the New York Flora Atlas only in Bronx and Staten Island (2008), are treated as a single species (R. bifrons) by the Flora of North America. However in Eurasia, they appear to be two distinct species. iNaturalist allows both species, following Kew's POWO, and both have a significant number of research-grade observations in North America. Rosales Rosaceae Rosoideae Rubeae Rubus Rubus Rubus Rubus Blackberry 843 30 Rubus Rubus Hiemales Rubus laciniatus Willd. Willdenow 1806 Rubus laciniatus Willd. 1822 Rubus corylifolius SM.? laciniatus (Willd.) Wallr. 1858 Rubus nemoralis P.J.Müll. (POWO accepted name 2023) 1893 Rubus vulgaris P.J.Müll. laciniatus (Weston) Dippel 1902 Rubus bifrons Vest laciniatus (Willd.) Murr 1909 Rubus bifrons Vest laciniatus (Weston) Dalla Torre & Sarnth. 1914 Rubus vulgaris P.J.Müll. laciniatus (Weston) Frid. 2014 Rubus nemoralis P.J.Müll. laciniatus (Willd.) A.Beek Cutleaf blackberry Cut-leaved blackberry Evergreen blackberry Cutleaf bramble Ronce laciniée Introduced Moderately invasive 63%Rubus laciniatus Moderate 63 all 6531 X Cayu RULA XX 4TKMN 2992543 78891-Rubus-laciniatus 738594-1 0000984132 8936 32364 242417164 Rubus_laciniatus Rubus in Poznan Lawica.jpg Rubus bifrons Vest ex Tratt. 1821 Rubus bifrons Vest 1823 Rubus bifrons Vest ex Tratt. 1825 Rubus linkianus Ser. 1846 Rubus nemorosus Hayne ex Willd. bifrons (Vest) Bab. 1874 Rubus armeniacus Focke (per FNA 2023) auct Rubus procerus non P.J.Müll. ex Genev 1898 Rubus fruticosus L. bifrons (Vest) Fiori Himalayan berry 2023 GBIF Rubus bifrons Vest ex Tratt. Himalayan blackberry 2023 New York Flora Atlas, Rubus bifrons Vest Himalayan giant blackberry 2023 VASCAN: GRIN Twice-leaved blackberry 2023 VASCAN: M.Anions Ronce discolore Introduced Europe Moderately invasive No NY specimens 57%Rubus bifrons Moderate 57 L 2754 X none RUBI X Bronx (only) 168025-Rubus-bifrons 60449596-2 8925 24891 32274 27801204 nymap.svg 2 counties 4 counties Rubus bifrons - Botanischer Garten, Frankfurt am Main - DSC02471.JPG Rubus armeniacus Focke Rubus bifrons Vest (per FNA 2023) 1874 Rubus armeniacus Focke 1906 Rubus macrostemon (Focke) Caflisch armeniacus (Focke) Sprib. 1914 Rubus hedycarpus Focke armeniacus Focke auct Rubus discolor non Weihe & Nees (1824) Armenian Blackberry Himalayan blackberry Himalaya giant blackberry Introduced Caucasus Moderately invasive IO 57%Rubus bifrons Moderate 57 L 0 0 Not listed RUAR9 X MA,NJ,PA,OH,ON 4TJGY 735202-1 837296 32263 nymap.svg Synonymized 1 county 20190624 Rubus armeniacus Drouwenerzand.jpeg Rubus ulmifolius Schott Erneuerte Vaterl. Blätt. Oesterr. Kaiserstaat 11: 42 (1818), nom. cons. prop. 1788 Rubus inermis Pourr. 1809 Rubus inermis Willd. 1818 Rubus ulmifolius Schott 1824 Rubus discolor Weihe & Nees 1825 Rubus flagellaris Willd. inermis Ser. 1861 Rubus rusticanus Mercier 1869 Rubus bellidiflorus hort. ex K.Koch 1903 Rubus ulmifolius Schott genuinus Sudre 1903 Rubus ulmifolius Schott vulgatus Sudre 1905 Rubus ulmifolius Schott bellidiflorus C. K. Schneid. 1914 Rubus ulmifolius Schott inermis (Ser.) Focke Elmleaf blackberry 2021 iNaturalist Elm-leaved blackberry 2021 NYFA Introduced Potentially invasive 7604 Xm 2996929 78895-Rubus-ulmifolius 740982-1 0000985000 313548 nymap.svg Bronx (2016) No RG observations Macchiabjörnbär (Rubus ulmifolius)-2975 - Flickr - Ragnhild & Neil Crawford.jpg Rubus subg. Rubus sect. Hispidi Rubus-hispidus-Acadia.jpg Rubus hispidus Rosales Rosaceae Rosoideae Rubeae Rubus Rubus Hispidi Rubus Dewberry 843 30 Rubus Hispidi Rubus hispidus L. 1753 Rubus hispidus L. 1923 Rubus hispidus L. blanchardianus L.H.Bailey 1824 Rubus sempervirens Bigelow 1940 Rubus hispidus L. obovalis Fernald 1941 Rubus blanchardianus (L.H.Bailey) L.H.Bailey 1941 Rubus fassettii L.H.Bailey 1941 Rubus hispidus L. cupulifer L.H.Bailey 1941 Rubus hispidus L. pervarius L.H.Bailey Bristly dewberry Running blackberry Swamp blackberry Swamp dewberry Ronce hispide Native Secure 2756 5 RUHI NN 8931 rubus/hispidus 24943 32340 27801261 Bristly Dewberry, Swamp Dewberry (Rubus hispidus) (2864363698).jpg Rubus subg. Rubus sect. Flagellares Rubus flagellaris UGA1120430.jpg Rubus flagellaris The Flora of North America Editorial Committee (2014) treats both the Arundel blackberry (R. arundelanus), Enslen's blackberry (R. enslenii), and the Ithaca blackberry (R. ithacanus) as being among the 112 synonyms of the American dewberry (R. flagellaris).Rubus flagellaris Willd. at Tropicos.org. Missouri Botanical Garden. 04 Jul 2015 <http://www.tropicos.org/Name/27801236>Rubus flagellaris Willd. at Canadensys, VASCAN last updated on 2015-06-29 Rosales Rosaceae Rosoideae Rubeae Rubus Rubus Flagellares Rubus Dewberry 843 30 Rubus Flagellares Rubus flagellaris Willd. Enum. Pl. Hort. Berol. 2: 549 (1809) 1809 Rubus flagellaris Willd. 1894 Rubus baileyanus Britton 1901 Rubus procumbens Rydb 1906 Rubus arenicola Blanch. 1906 Rubus geophilus Blanch American dewberry Northern dewberry Whiplash dewberry Northern blackberry Ronce à flagelles Native Secure 2757 5 RUFL NN 8929 rubus/flagellaris 24921 104958 242417161 27801236 Rubus flagellaris WFNY-093B.jpg Rubus subg. Rubus sect. Caesii Rubus caesius SSN-150707-01.jpg Rubus caesiusEurasian dewberry The European dewberry has only been reported outside of cultivation in limited number of counties in North America, including only a few from New York. As of July 2015, it was not listed on New York State's Non-Native Plant Species Invasiveness Assessment.Non-Native Plant Species Invasiveness Assessment However the Finger Lakes Native Plant Society,Rubus caesius: Finger Lakes Native Plant Society website 2015. Cornell Plantations, and Tompkins CountyTompkins County 2009 Regional Invasive Species List all list Rubus caesius as an invasive plant. Rosales Rosaceae Rosoideae Rubeae Rubus Rubus Caesii Rubus Dewberry 843 30 Rubus Caesii Rubus caesius L. 1753 Rubus caesius L. 1782 Rubus caeruleus Gilib. 1829 Rubus polymorphus caesius (L.) Spenn. 1888 Rubus caesius intermedia Schmidely 1889 Rubus caesius L. eucaesius Bouvet (not validly publ.) 1891 Rubus caesius turkestanicus Regel 1894 Rubus fruticosus L. caesius Bonnier & Layens 1895 Selnorition cesius (L.) Raf. ex B.D.Jacks. 1898 Rubus fruticosus L. caesius (L.) Fiori 1937 Rubus psilophyllus Nevski (nom. illeg.) Eurasian dewberry European dewberry Ronce bleue Ronce bleuâtre Introduced Eurasia northern Africa Moderately invasive CP-3CP3 2753 X Onon RUCA XX CP-3CP3 2995209 735608-1 8926 0 24898 32278 27800345 Rubus%20caesius nymap.svg 5 counties 11 counties Gervuoge001.jpg Rubus subg. Cylactis Rubus pubescens flower.jpg Rubus pubescensdwarf raspberry Rosales Rosaceae Rosoideae Rubeae Rubus Cylactis Rubus Blackberry 843 30 Cylactis Rubus pubescens Raf. Med. Repos. 2: 333 (1811) 1803 Rubus saxatilis Michx. (nom. illeg.) 1811 Rubus pubescens Raf. 1819 Cylactis montana Raf. 1823 Rubus triflorus Richardson 1824 Rubus canadensis Torr. (sensu auct.) 1825 Rubus mucronatus Ser. 1893 Cylactis egopodioides (Ser.) Raf. ex B.D.Jacks. 1985 Cylactis pubescens (Raf.) W.A.Weber Dwarf raspberry Dwarf red blackberry Creeping red raspberry Trailing red raspberry Plumboy Ronce pubescente Catherinette Native Secure 8 FACW Perennial Shrub S5 6534 5 RUPUP2 NN 284431-2 8945 rubus/pubescens 25028 32425 250100444 27800180 Rubus pubescens 1-eheep (5097346393).jpg Tribe Colurieae Waldsteinia fragarioides WFNY-099.jpg Appalachian barren strawberry The New York members of the Colurieae tribe include about ten species of the genus Geum (avens) and a single species of Waldsteinia (barren strawberry). Modern phylogenetic evidence, including studies by Protopopova et al. (2022, 2023), supports treating Waldsteinia fragarioides as a separate genus rather than combining it under Geum M V Protopopova et al 2021 IOP Conf. Ser.: Earth Environ. Sci. 908 012027 DOI 10.1088/1755-1315/908/1/012027Protopopova, Marina, et al. 2023. "Waldsteinia within Geum s.l. (Rosaceae): Main Aspects of Phylogeny and Speciation History" Diversity 15, no. 4: 479. https://doi.org/10.3390/d15040479. Full text and detailed descriptions can be found in the referenced web documents . Waldsteinia Barren Strawberry (Geum fragarioides) - Guelph, Ontario 01.jpg Waldsteinia fragarioides The type species of the genus Waldsteinia was historically nested within Geum by J. E. E. Smedmark in 2006 James B. Phipps (2020). FNA Volume 9. page 71. Flora of North America Association 5 Nov. 2020, but multi-gene sequencing has since reinstated its distinct genus status Weakley, A.S., and Southeastern Flora Team (2023). Flora of the southeastern United States Web App.Protopopova, Marina, et al. (2023). "Waldsteinia within Geum s.l. (Rosaceae)". Rosales Rosaceae Rosoideae Colurieae Waldsteinia Waldsteinia Barren strawberry 886 10 Waldsteinia fragarioides (Michx.) Tratt. fragarioides 1803 Dalibarda fragarioides Michx. Fl. Bor.-Amer. 1: 300 (1803) 1814 Rubus fragarioides (Michx.) Dum.Cours. Bot. Cult., ed. 2, 7: 293 (1814) 1823 Waldsteinia fragarioides (Michx.) Tratt. Rosac. Monogr. 3: 107 (1823) 1825 Comaropsis fragarioides (Michx.) DC. ex Ser. A.P.de Candolle, Prodr. 2: 555 (1825) 1825 Dryas trifoliata Pall. ex Ser. A.P.de Candolle, Prodr. 2: 555 (1825) 1840 Bossekia fragarioides (Michx.) Raf Autik. Bot.: 175 (1840) 2006 Geum fragarioides (Michx.) Smedmark Bot. Jahrb. Syst. 126: 415 (2006) synonym layout 2023 Waldsteinia fragarioides (Michx.) Tratt. reinstated by Protopopova et al. acc. by POW, Weakley Appalachian barren strawberry Northern Barren Strawberry Dry strawberry Waldsteinie faux-fraisier Native Secure 7 Perennial Herb-forb S5, G5 2686 5 Waldsteinia fragarioides WAFR NN Waldsteinia fragarioides 6KFTJ 5370015 1168107-2 3122 8990 geum/fragarioides 894458-Waldsteinia-fragarioides 505735 464850 242417492 Waldsteinia_fragarioides 50326515 nymap.svg Waldsteinia fragarioides 15 (5097400745).jpg Geum Geum canadense 2533.jpg Geum canadense The New York Flora Atlas lists ten species of Geum, including Geum fragarioides, which is here treated above as Waldsteinia fragarioides. Rosales Rosaceae Rosoideae Colurieae Geum Geum Avens 886 10 L. Geum canadense Jacq. Jacquin, Nicolaus(Nicolaas) Joseph von (1727-1817) Hortus Botanicus Vindobonensis (Hort. Bot. Vindob.) 1773 Geum canadense Jacq. 1775 Geum laciniosum Murray 1788 Geum carolinianum Walter 1791 Geum album J.F.Gmel. 1794 Caryophyllata alba Moench 1831 Geum glutinosum Pursh ex Lehm. 1832 Sieversia caroliniana G.Don 1913 Geum camporum Rydb 1922 Geum canadense Jacq. camporum (Rydb.) Fernald & Weath. 1922 Geum canadense Jacq. grimesii Fernald & Weath. 1922 Geum canadense Jacq. texanum Fernald & Weath. 1937 Geum canadense Jacq. brevipes Fernald White avens Canada avens Benoîte du Canada Benoîte blanche Native Secure 5 FAC FACU Perennial Herb-forb S5, G5 2684 5 GECA7 NN 3FY96 5369800 85325-Geum-canadense 30065568-2 0001012938 3120 8765 geum/canadense 316698 242416592 Geum_canadense 27800887 30065568-2 Geum+canadense Geum%20canadense nymap.svg Geum canadense 2826.jpg Geum aleppicum Jacq. 1781 Geum aleppicum Jacq. 1789 Geum strictum Aiton (nom. superfl.) 1913 Geum decurrens Rydb. 1824 Geum ranunculoides Ser. 1829 Geum rugosum Desf. 1935 Geum aleppicum Jacq. strictum (Aiton) Fernald 1939 Geum strictum Aiton decurrens (Rydb.) Kearney & Peebles 1949 Geum aleppicum Jacq. strictum (Aiton) R.T.Clausen Yellow avens Aleppo avens Common avens Benoîte d’Alep Native Secure 3 6509 5 GEAL3 NN 30094401-2 8763 geum/aleppicum 17514 200010864 27800066 nymap.svg Yellow Avens (Geum aleppicum) (5748647868).jpg Geum rivale L. 1753 Geum rivale L. 1771 Caryophyllata rivalis (L.) Scop. 1790 Anemone decaphilla Krock. 1790 Anemone dodecaphilla Krock. 1791 Anemone decapetala J.F.Gmel. 1840 Bernoullia hybrida Raf. 1840 Bernoullia nutans (Raf.) Raf. 1840 Bernoullia rivalis (L.) Raf. Purple avens Water avens Chocolate-root Benoîte des ruisseaux Benoîte pourpre Native Secure 8 OBL 2689 5 GERI2 NN 725355-1 8772 geum/rivale 200010867 nymap.svg Geum rivale 2015-06-01 OB 110.jpg Geum laciniatum Murray Novi Comment. Soc. Regiae Sci. Gott. 5: 30 (1775) 1775 Geum laciniatum Murray 1935 Geum laciniatum Murray trichocarpum Fernald Rough avens Floodplain avens Benoîte lacinicée Native Secure 3 FACW FAC Perennial Herb-forb S5, G5 2683 5 GELA NN 3FYBG 5369645 163272-Geum-laciniatum 725280-1 0001012963 8767 geum/laciniatum 242416593 Geum_laciniatum nymap.svg Geum laciniatum var. trichocarpum ROUGH AVENS (3328625944).jpg Geum vernum (Raf.) Torr. & A.Gray Fl. N. Amer. 1: 422 (1840) 1821 Stylypus vernus Raf. 1821 Stypostylis vernus Raf. 1840 Geum vernum (Raf.) Torr. & A.Gray Spring avens N. America native NY nativity uncertain 5 FACU Perennial Herb-forb SNR, G5 2648 16 counties 16 counties 130906 11 counties 11 counties 5369957 present in NY present in New York State GEVE NN Endangered in NY 725402-1 geum/vernum Geum_vernum nymap.svg Spring Avens (Geum vernum) - Flickr - Jay Sturner (1).jpg Geum triflorum Pursh triflorum 1814 Geum triflorum Pursh 1823 Sieversia triflora (Pursh) R.Br. 1832 Geum pubescens Hook. 1906 Erythrocoma campanulata (Greene 1906 Erythrocoma canescens (Greene 1906 Erythrocoma flavula (Greene 1906 Erythrocoma triflora (Pursh) Greene 1936 Geum ciliatum Pursh triflorum (Pursh) Jeps. (nom. illeg.) 1958 Geum canescens (Greene) Munz Prairie smoke Old man's whiskers Native Threatened 10 FACU Perennial Herb-forb S2, G5 2666 2 GETR NN 109871-2 0 nymap.svg The botanical cabinet (No 1609) (8613654108).jpg Geum virginianum L. 1753 Geum virginianum L. 1785 Caryophyllata virginiana (L.) Lam. 1813 Geum hirsutum Muhl. ex Link 1822 Geum heterophyllum Link (nom. nud.) 1828 Geum virginianum L. lanceolatum Raf. (nom. nud.) 1828 Geum virginianum L. macrophyllum Raf. (nom. nud.) 1828 Geum virginianum L. ochroleucum Raf. (nom. nud.) 1828 Geum virginianum L. ramosum Raf. (nom. nud.) 1828 Geum virginianum L. uniflorum Raf. (nom. nud.) 1889 Geum album J.F.Gmel. flavum Porter 1896 Geum flavum (Porter) E.P.Bicknell 1923 Geum virginianum L. murrayanum Fernald Cream-colored avens Cream avens Native Threatened 9 S2 FACU FAC Perennial Herb-forb S2, G5 6511 23 counties 23 counties 131001 Livingston, Wyoming Livingston, Wyoming 5369702 present in NY present in New York State GEVI4 NN geum/virginianum 24665 nymap.svg Geum virginianum iNat-146376349 (2x3).jpg Geum macrophyllum Willd. Enum. Pl. Hort. Berol. 2: 557 (1809) macrophyllum 1809 Geum macrophyllum Willd. 1910 Geum fauriei H.Lév. 1915 Geum japonicum Thunb. sachalinense Koidz. 1924 Geum japonicum Thunb. fauriei (H.Lév.) Kudô 1926 Geum sachalinense (Koidz.) Makino 1952 Geum macrophyllum Willd. sachalinense (Koidz.) H.Hara 1953 Geum aleppicum Jacq. sachalinense (Koidz.) Ohwi 1967 Geum perincisum Rydb. intermedium B.Boivin 1982 Geum macrophyllum Willd. fauriei (H.Lév.) Vorosch. Largeleaf avens Large-leaved avens Native Endangered 8 S1 FACW Perennial Herb-forb S1, G5 2663 10 counties Chenango, Essex, Orange, Orleans, Otsego, Saratoga, Steuben, Ulster, Warren, Yates 77266 Delaware, Ulster Delaware, Ulster Counties 5369784 present in NY present in New York State GEMAM NN geum/macrophyllum nymap.svg Geum macrophyllum 202912270.jpg Geum urbanum L. 1753 Geum urbanum L. 1771 Caryophyllata urbana (L.) Scop. 1794 Caryophyllata officinalis Moench 1840 Bernoullia media Raf. 1893 Geranium pseudomolle (Pant.) B.D.Jacks. 1974 Geum rivale L. urbanum (L.) Á.Löve & D.Löve Town avens Herb Bennet Bennet's root Wood avens Clove-root Colewort St. Benedict's herb Benoîte commune Introduced Invasive 2725 7 counties Cayuga (2006), Lewis (2026), Monroe (1989), Onondaga (1988), Orange (1990), St. Lawrence (1997), Yates (2021) 55891-Geum-urbanum 22 counties GEUR XX 8776 geum/urbanum 24663 17518 Geum%20urbanum Geum urbanum nymap.svg 20130617Geum urbanum.jpg Geum hybrids Geum xcatlingii comparison.jpg Geum × catlingii (center),Geum canadense (top right),Geum urbanum (bottom) Rosales Rosaceae Rosoideae Colurieae Geum Geum Avens 886 10 L. Geum × aurantiacum Fr. ex Scheutz Prodr. Monogr. Geor.: 30 (1870) Geum aleppicum Geum rivale 1870. Geum × aurantiacum Fr. ex Scheutz in Prodr. Monogr. Geor.:30 Orange avens Hybrid yellow & purple avens Hybrid yellow & water avens Native Unranked Perennial Herb-forb SNA, GNA 6510 Onondaga (1942) Onondaga (1942, M.E. Faust & N.M. Sadler) 170751 no NY observations 0 5370010 not present in NY not present in New York State GEAU2 NN 8V8DS 725183-1 geum/aleppicum 24648 nymap.svg Ny hybrid.svg Geum × cortlandicum M.Hough Phytoneuron 2018-59: 1 (2018) Geum canadense Geum laciniatum 2018. Geum × cortlandicum M.Hough in Phytoneuron 2018-59:1 Cortland avens Hybrid white & rough avens Native Unranked SNA, GNA 7655 Cortland, Seneca Cortland (2018), Seneca (2018) 954291 Cortland, Seneca Cortland (2018, 2020), Seneca (2018) 10716832 Cortland, Seneca Cortland (2018, 2020), Seneca (2018) 6KFVW 77203992-1 nymap.svg Ny hybrid.svg Geum × hainesianum M.Hough, A.V.Gilman & Chapm.-Lam 2021. Geum x hainesianum (Rosaceae), a new hybrid avens from eastern North America. Rhodora 123(993): 67-84. Geum aleppicum Geum canadense 2022. Geum × hainesianum M.Hough, A.V.Gilman & Chapm.-Lam in Rhodora 123: 13 Haines’s avens Hybrid yellow & white avens Native Unranked SNA, GNA 7613 Tioga (1942) Tioga (1942, R.T. & E.R.Clausen) 1426000 Cortland (cultivated) Cortland (2021, cultivated by M. Hough) 12231213 not present in NY not present in New York State B2Y5W 77309022-1 nymap.svg Ny hybrid.svg Geum × catlingii J.-P.Bernard & R.Gauthier Naturaliste Canad. 113: 321 (1986) Geum canadense Geum urbanum 1986. Geum × catlingii J.-P.Bernard & R.Gauthier in Naturaliste Canad. 113:321 Catling's avens Hybrid white & town avens Introduced Naturalized Perennial Herb-forb SNA, GNA 7614 Cortland (2016) Cortland (2016, M. Hough) 170752 9 NY counties Cayuga, Chautauqua, Chemung, Cortland, Erie, Monroe, Onondaga, Schuyler, Tompkins 5369947 present in NY present in New York State GECA17 N? BS9BV 931308-1 nymap.svg Geum xcatlingii fl.jpg Geum × intermedium Ehrh. Beitr. Naturk. Verw. Wiss. 6: 143 (1791) Geum rivale Geum urbanum 1791. Geum × intermedium Ehrh. in Beitr. Naturk. Verw. Wiss. 6:143 1800. Geum urbanum intermedium (Ehrh.) Willd. in Sp. Pl., ed. 4. 2: 1114 1812. Geum urbanum var. intermedium (Ehrh.) Mérat in Nouv. Fl. Env. Paris:198 1825. Geum rivale var. intermedium (Ehrh.) Ser. in A.P.de Candolle, Prodr. 2:551 1834. Geum rivale ssp. intermedium (Ehrh.) Schübl. & G. Martens 1836. Geum × rivale-urbanum G.Mey. in Chloris Han.:39 (not validly publ.) 1841. Geum × willdenowii J.N.Buek bis in Fl. March.:595 1860. Geum × intermedium var. willdenowii (J.N.Buek bis) Asch. in Fl. Brandenburg 1:179 Intermediate avens Hybrid purple & town avens Introduced Naturalized Perennial Herb-forb SNA, GNA 7656 Cortland (2018) Cortland (2018, M. Hough) 628500 Cayuga, Cortland Cayuga (2018, 2021), Cortland (2018-2023), Madison (2023, not RG yet) 5369692 Cayuga, Cortland Cayuga (2018, 2021), Cortland (2018-2023) 7VQST 725266-1 nymap.svg Geum × intermedium 002.jpg Geum × macneillii J.-P.Bernard & R.Gauthier Naturaliste Canad. 113: 321 (1986) Geum laciniatum Geum urbanum 1986. Geum × macneillii J.-P.Bernard & R.Gauthier in Naturaliste Canad. 113:321 McNeill's avens Hybrid rough & town avens Introduced Naturalized Perennial Herb-forb SNA, GNA 7657 Cortland (2018) Cortland (2018, M.Hough) 975912 Cayuga, Cortland Cayuga (2021), Cortland (2019-22) 5369940 Cayuga, Cortland Cayuga (2021), Cortland (2019-22) CDBGY 931307-1 nymap.svg Ny hybrid.svg