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'The All-Species Living Tree' Project
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Collaborative academic project
February 2021
June 2025
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The All-Species Living Tree' Project (LTP) is a collaboration between various academic groups/institutes, such as ARB, SILVA rRNA database project, and LPSN, with the aim of assembling a database of 16S rRNA sequences of all validly published species of Bacteria and Archaea. Yarza P. Richter M. Peplies J. R. Euzeby J. Amann R. Schleifer K. H. Ludwig W. Glöckner F. O. Rosselló-Móra R. 10.1016/j.syapm.2008.07.001 The All-Species Living Tree project: A 16S rRNA-based phylogenetic tree of all sequenced type strains Systematic and Applied Microbiology 31 4 241–250 2008 18692976 2008SyApM..31..241Y 10261/103580 https://tede.ufrrj.br/jspui/handle/jspui/5297 free At one stage, 23S sequences were also collected, Yarza
Pablo
Ludwig
Wolfgang
Euzéby
Jean
Amann
Rudolf
Schleifer
Karl-Heinz
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Frank Oliver
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Update of the All-Species Living Tree Project based on 16S and 23S rRNA sequence analyses
Systematic and Applied Microbiology
33
6
291–299
2010
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10.1016/j.syapm.2010.08.001 2010SyApM..33..291Y
but this has since stopped. Munoz R. L. Yarza P. Ludwig W. Euzéby J. Amann R. Schleifer K. H. Oliver Glöckner F. Rosselló-Móra R. 10.1016/j.syapm.2011.03.001 Release LTPs104 of the All-Species Living Tree Systematic and Applied Microbiology 34 3 169–170 2011 21497273 2011SyApM..34..169M
Currently there are over 10,950 species in the aligned dataset and several more are being added either as new species are discovered or species that are not represented in the database are sequenced. Initially the latter group consisted of 7% of species.
Similar (and more recent) projects include the Genomic Encyclopedia of Bacteria and Archaea (GEBA), which focused on whole genome sequencing of bacteria and archaea.Wu Dongying Hugenholtz Philip Mavromatis Konstantinos Pukall Rüdiger Dalin Eileen Ivanova Natalia N. Kunin Victor Goodwin Lynne Wu Martin Tindall Brian J. Hooper Sean D. December 2009 A phylogeny-driven genomic encyclopaedia of Bacteria and Archaea Nature 462 7276 1056–1060 10.1038/nature08656 0028-0836 3073058 20033048 2009Natur.462.1056WKyrpides Nikos C. Hugenholtz Philip Eisen Jonathan A. Woyke Tanja Göker Markus Parker Charles T. Amann Rudolf Beck Brian J. Chain Patrick S. G. Chun Jongsik Colwell Rita R. 5 August 2014 Genomic Encyclopedia of Bacteria and Archaea: Sequencing a Myriad of Type Strains PLOS Biology 12 8 e1001920 10.1371/journal.pbio.1001920 1545-7885 4122341 25093819 free
Tree
Bacterial phyla
The tree was created by maximum likelihood analysis without bootstrap: consequently accuracy is traded off for size and many phylum level clades are not correctly resolved (such as the Firmicutes). (Eukaryotes not present in analysis). This phylogeny is a summary of the 16S rRNA based LTP_07_2026 and contains all type species with validly published names up to July 2026.The LTPlus https://bioinfo.uib.es/ltp/#LTPlus The All-Species Living Tree Project 1 August 2026LTP_all tree in newick format https://biocom.uib.es/opucheck-backend/api/releases/07_26_04 The All-Species Living Tree Project 1 August 2026Rossello-Mora Ramon Viver Tomeu Palmer-Rodríguez Pere Westram Ralf Bustos-Caparros Esteban Gago Joan Francisco Gonzalez-Mendez Jaime Llabrés Mercè Ludwig Wolfgang Amann Rudolf A pipeline for improved 16S rRNA gene-based phylogeny and diversity analyses of Bacteria and Archaea Nature Portfolio 22 April 2026 10.21203/rs.3.rs-9370187/v1 https://doi.org/10.21203/rs.3.rs-9370187/v1 en free
font-size:90%;line-height:90% Domain Archaea
Microcaldota
Methanopyri
Nanobdellati
Nanobdellota
Promethearchaeati
Promethearchaeota
Thermoproteati
Thermoproteota [incl. Nitrososphaeria; Methanosuratincolia]
Methanobacteriati
Methanobacteria
Thermococci
Methanococci
"Archaeoglobia"
Thermoplasmata
Methanonatronarchaeia
Methanocellia
Methanosarcinia
Methanomicrobia
Halobacteria
Domain Bacteria
Minisyncoccota
Bacteriovoracia
Abditibacteriota
Armatimonadota
Oligoflexia
Thermodesulfobiota
Thermotogota
Bradymonadia
"Fervidibacterota"
Thermoflexia
Coprothermobacterota
Aquificota
Syntrophorhabdia
Atribacterota
Elusimicrobiota
Deinococcota
Brevinematia
Myxococcota
"Desulfuromonadota"
Cyanobacteriota [incl. Vampirovibrionia]
Thermodesulfovibrionia
Bacillota [incl. Desulfuribacillia; Erysipelotrichia]
"Desulfatigladia"
"Leptospirillaeota"
Acidobacteriota
Polyangiia
Chlamydiota
Kiritimatiellota
Lentisphaerota
Verrucomicrobiota
Desulfomonilia
Syntrophia
Chloroflexota
Desulfobacterota
Planctomycetota
Actinomycetota
Bdellovibrionota
Thermosulfidibacterota
Thermodesulfobacteriota
"Desulfovibrionota"
Deltaproteobacteria~ [incl. Desulfobaccia; Dissulfuribacteria; Desulfarculia; Syntrophobacteria; Desulfobulbia]
"Clostridiota"
Chlorobiota [incl. Ignavibacteria]
Balneolota
Rhodothermota
Bacteroidota
Caldisericota
Vulcanimicrobiota
Gemmatimonadota
Fusobacteriota
Synergistota
Spirochaetota
Deferrisomatota
Chrysiogenota
Deferribacterota
Campylobacterota
"Alphaproteobacteriota"
Fidelibacterota
Calditrichota
Fibrobacterota
Chromatibacteria [incl. Betaproteobacteria, Gammproteobacteria, Zetaproteobacteria, Magnetococcia]
See also
Branching order of bacterial phyla (Woese, 1987)
Branching order of bacterial phyla (Gupta, 2001)
Branching order of bacterial phyla (Cavalier-Smith, 2002)
Branching order of bacterial phyla (Rappe and Giovanoni, 2003)
Branching order of bacterial phyla (Battistuzzi et al., 2004)
Branching order of bacterial phyla (Ciccarelli et al., 2006)
Branching order of bacterial phyla (Genome Taxonomy Database, 2018)
Bacterial phyla
List of Archaea genera
List of bacteria genera
List of bacterial orders
LPSN, list of accepted bacterial and archaeal names
References
External links
http://www.arb-silva.de/projects/living-tree/
Category:Bacteria
Category:Bacteriology
Category:Microbiology
Category:Taxonomy (biology)
Category:Systems of bacterial taxonomy