411 REFERENCE · Wikipedia
1999 in paleontology
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1999 paleontology paleobotany arthropod paleontology paleoentomology paleomalacology reptile paleontology archosaur paleontology mammal paleontology paleoichthyology
1999
Flora
Ferns and fern allies
NameNoveltyStatusAuthorsAgeType localityLocationNotesImagesMakotopterisStockey R. A. Nishida H. Rothwell G. W. 1999 Permineralized ferns from the middle Eocene Princeton chert. I. Makotopteris princetonensis gen. et sp. nov.(Athyriaceae) International Journal of Plant Sciences 160 5 1047–1055 10.1086/314191 10506480 1999IJPlS.160.1047S 33465214 Gen et sp novvalidStockey, Nishida, & RothwellEoceneYpresianEocene Okanagan HighlandsAllenby FormationPrinceton ChertCanadaBritish ColumbiaAn athyriaceous fern The type species is M. princetonensis
Cycads
NameNoveltyStatusAuthorsAgeType localityLocationNotesImagesEostangeria pseudopterisKvaček Z. Manchester S.R. 1999-05-01 Eostangeria Barthel (Extinct Cycadales) from the Paleogene of Western North America and Europe International Journal of Plant Sciences 160 3 621–629 10.1086/314152 10.1086/314152 1999IJPlS.160..621K 83978433 1058-5893Sp novvalidKvaček & ManchesterLate Paleoceneupper Fort Union Formation or lower Wasatch FormationUSAWyomingAn eostangerioid zamiaceous cycad.Also found in the Eocene John Day Formation of Oregon.
Angiosperms
NameNoveltyStatusAuthorsAgeType localityLocationNotesImagesFlorissantia sikhote-alinensisManchester S. R. 1999 Biogeographical relationships of North American tertiary floras Annals of the Missouri Botanical Garden 86 2 472–522 10.2307/2666183 2666183 1999AnMBG..86..472M https://www.biodiversitylibrary.org/part/28623 comb novvalid(Kryshtofovich) ManchesterMioceneAmgu floraRussiaPrimorsky KraiA mallow relative, moved from Porana sikhote-alinensis (1921) Quercus hiholensisBorgardt S. J. Pigg K. B. 1999 Anatomical and developmental study of petrified Quercus (Fagaceae) fruits from the Middle Miocene, Yakima Canyon, Washington, USA American Journal of Botany 86 3 307–325 10.2307/2656753 2656753 10077494 1999AmJB...86..307B free Sp novValidBorgardt & PiggMiddle Miocene"Yakima Canyon Flora"USAA white oak, permineralized acornsthumb|center|upright|†Quercus hiholensisSargentodoxa globosaComb novvalid(Manchester) ManchesterMiddle EoceneClarno FormationUSAOregonA lardizabalaceous species; New comb for Bumelia? globosa (1994)
Arthropods
Insects
Name Novelty Status Authors Age Unit Location Notes ImagesCephalotes alveolatusde Andrade M. L. Baroni Urbani C. 1999 Diversity and adaptation in the ant genus Cephalotes, past and present Stuttgarter Beiträge zur Naturkunde, Serie B (Geologie und Paläontologie) 271 537–538Comb novValid(Vierbergen & Scheven)BurdigalianDominican amberDominican RepublicA myrmicinae ant, Moved from Zacryptocerus alveolatus (1995)thumb|center|upright|Cephalotes alveolatusCephalotes caribicusSp novValidDe Andrade & Baroni UrbaniBurdigalianDominican amberDominican RepublicA myrmicinae ant.thumb|center|upright|Cephalotes caribicusCephalotes dieteriSp novValidDe Andrade & Baroni UrbaniBurdigalianDominican amberDominican RepublicA myrmicinae ant.thumb|center|upright|Cephalotes dieteriCephalotes hispaniolicusSp novValidDe Andrade & Baroni UrbaniBurdigalianDominican amberDominican RepublicA myrmicinae ant.thumb|center|upright|Cephalotes hispaniolicusCephalotes serratusComb novvalid(Vierbergen & Scheven, 1995)BurdigalianDominican amberDominican RepublicA myrmicinae ant, moved from Exocryptocerus serratus (1995)Nine new species and a new genus of Dominican amber ants of the tribe (Cephalotini Hymenoptera: Formicidae) Creation Research Society Quarterly Vierbergen GS, Scheven J 31 December 1995 27 March 1995 AntWiki 158–170 https://www.antwiki.org/wiki/images/1/15/Vierbergen_&_Scheven_1995.pdf thumb|center|upright|Cephalotes serratusCamponotus crozeiRiou B. 1999 Descriptions de quelques insectes fossiles du Miocène supérieur de la Montagne d'Andance (Ardèche, France) Travaux de l'École Pratique des Hautes Études, Biologie et Évolution des Insectes 11/12 123–133Sp novValidRiouTurolianMontagne d'AndanceFranceA formicine ant.thumb|center|upright|Camponotus crozeiElectrostephanus neovenatusEngel M.S. Ortega-Blanco J. 2008 The fossil crown wasp Electrostephanus petiolatus Brues in Baltic Amber (Hymenoptera, Stephanidae): designation of a neotype, revised classification, and a key to amber Stephanidae ZooKeys 4 55–64 10.3897/zookeys.4.49 2008ZooK....4...55E free 2445/36428 free Sp novvalidAguiar & JanzenLutetianBaltic AmberEuropeA stephanid waspElectrostephanus sulcatusSp novjr synonymAguiar & JanzenLutetianBaltic AmberEuropeA stephanid wasp, Moved to Denaeostephanus sulcatusNeoephemera antiquaSinitchenkova N. D. A new mayfly species of the extant genus Neoephemera from the Eocene of North America (Insecta: Ephemerida=Ephemeroptera) Paleontological Journal 1999 33 4 403–405sp. novvalidSinitchenkovaYpresian
Klondike Mountain FormationUSAA neoephemerid mayflythumb|center|upright|Neoephemera antiquaPachycondyla rebekkaeRust J. Andersen N. M. Giant ants from the Paleogene of Denmark with a discussion of the fossil history and early evolution of ants (Hymenoptera: Formicidae) Zoological Journal of the Linnean Society 1999 125 3 331–348 10.1111/j.1096-3642.1999.tb00596.x 85900450 free sp. novvalidRust & AndersenYpresian
Fur FormationDenmarkA Myrmeciinae ant moved to Ypresiomyrma rebekkae (2006)Archibald S.B. Cover S. P. Moreau C. S. Bulldog Ants of the Eocene Okanagan Highlands and History of the Subfamily (Hymenoptera: Formicidae: Myrmeciinae) Annals of the Entomological Society of America 2006 99 3 487–523 http://antbase.org/ants/publications/21067/21067.pdf 10.1603/0013-8746(2006)99[487:BAOTEO]2.0.CO;2 4845957 thumb|center|upright|Pachycondyla rebekkaePenthetria intermediaFitzgerald S. J. 1999 A new species of Plecia from the Green River Formation and new combinations of fossil Bibionidae (Diptera). Great Basin Naturalist 59 182–187 10.5962/bhl.part.15821 free Comb novvalid(Scudder)PriabonianFlorissant FormationUnited StatesColorado
A bibionid fly Originally Mycetophaetus intermedius (1892)Scudder S. H. 1892 Some Insects of special interest from Florissant, Colorado and other points in the Tertiaries of Colorado and Utah Bulletin of the United States Geological Survey 93 1–25 Moved from Plecia intermedia (1959)Rice H. M. A 1959 Fossil Bibionidae (Diptera) from British Columbia Geological Survey of Canada Bulletin 55 1–36
Mollusca
Newly named bivalves
Name Novelty Status Authors Age Unit Location Notes ImagesConcavodonta ovalisSánchez T.M. 1999 New Late Ordovician (Early Caradoc) Bivalves from the Sierra de Villicum (Argentine Precordillera) Journal of Paleontology 73 1 66–76 1306745 10.1017/S0022336000027554 1999JPal...73...66S 133414164 sp novValid
SánchezCaradocDon Braulio FormationArgentinathird species in the genusConcavodontinaesubfam novValid
SánchezOrdovicianone of two Subfamilies in Praenuculidae CuyopsisGen et sp novValid
SánchezCaradocDon Braulio FormationArgentinaMonotypic with the species C. symmetricusEmilianiaGen et sp novhomonym
SánchezCaradocDon Braulio FormationArgentinagenus name preoccupied by Emiliania Hay & Mohler, 1967, renamed to EmiliodontaHemiconcavodontaGen et sp novValid
SánchezCaradocDon Braulio FormationArgentinathird genus in the subfamily ConcavodontinaePraenuculinaesubfam novValid
SánchezOrdovicianone of two Subfamilies in Praenuculidae Similodonta ceryxCope J.C.W. 1999 Middle Ordovician bivalves from Mid-Wales and the Welsh Borderland Palaeontology 42 3 467–499 10.1111/1475-4983.00081 1999Palgy..42..467C free Sp novValid
CopeMiddle OrdovicianGilfach Farm No. 1 BoreholeWalesoldest species in the genus SimilodontaTrigonoconchaGen et sp novValid
SánchezCaradocDon Braulio FormationArgentinaMonotypic with the species T. acutaVillicumiaGen et sp novValid
SánchezCaradocDon Braulio FormationArgentinaMonotypic with the species V. canteraensis
Fish
Newly named actinopterygii ("ray-finned fish")
Name Novelty Status Authors Age Unit Location Notes ImagesNardoveliferSorbini, C. Sorbini, L. The Cretaceous fishes of Nardo. 10: Nardovelifer altipinnis, gen. et sp. nov. (Teleostei, Lampridiformes, Veliferidae) St Ric Giac Terz Bolca 1999 8 11–27gen et sp novValidC. Sorbini & L. SorbiniCampanianNardoItalythumb|150px|center|Nardovelifer altipinnis
Reptiles
Archosauromorphs
Ornithomimid gastroliths documented.Kobayashi et al. (1999). Sanders, Manley, and Carpenter (2001), "Table 12.1" page 167.
Newly named dinosauriforms
Name Novelty Status Authors Age Unit Location Notes ImagesEucoelophysis Sullivan R.M. Lucas S.G. 1999 Eucoelophysis baldwini, a new theropod dinosaur from the Upper Triassic of New Mexico, and the status of the original types of Coelophysis Journal of Vertebrate Paleontology 19 1 81–90 10.1080/02724634.1999.10011124 1999JVPal..19...81S gen et sp novValid
Sullivan & LucasNorianChinle Formation, New MexicoUSASister genus to the Dinosaurs; first identified as a relative of coelophysid.
Newly named dinosaurs
Data courtesy of George Olshevsky's dinosaur genera list. Olshevsky George Dinogeorge's Dinosaur Genera List http://www.polychora.com/dinolist.html 2008-08-07 2011-07-15 https://web.archive.org/web/20110715102453/http://www.polychora.com/dinolist.html dead
Name Novelty Status Authors Age Unit Location Notes ImagesAchillobatorPerle, A., M. Norell and J. Clark. (1999). A new maniraptoran theropod Achillobator giganticus (Dromaeosauridae) from the Upper Cretaceous of Burkhant, Mongolia. Contributions of the Department of Geology, National University Mongolia (35 pages).Gen et sp novValidPerle, Norell, & ClarkLate CretaceousBayan Shireh FormationMongoliaA large dromaeosaurid.thumb|center|150px|AchillobatorAgustiniaBonaparte, J.F. (1999). An armored sauropod from the Aptian of Northern Patagonia, Argentina. Proceedings of the Second Gondwanan Dinosaur Symposium (15 October). National Science Museum Monographs 15 (Y. Tomida, T.H. Rich and P. Vickers-Rich, eds.), Tokyo.Gen et sp novValidBonaparteAlbianLohan Cura FormationArgentinaAn armored sauropodthumb|center|150px|Agustinia"Airakoraptor"gen novNomen nudum
Perle, Norell, & ClarkMongoliaA dromaeosaurid, now Kuru kulla, which was later described in 2021.AnimantarxCarpenter, K., K. Kirkland, D. Burge, and J. Bird. (1999). "Ankylosaurs (Dinosauria; Ornithischia) of the Cedar Mountain Formation, Utah, and the stratigraphic distributions". In: Vertebrate Paleontology in Utah (D. Gillete, ed.). Utah Geological Survey Miscellaneous Publication 99-1.Gen et sp novValidCarpenter, Kirkland, Burge, & BirdLate CretaceousCedar Mountain Formation, UtahUSANodosaur whose name means "living citadel."thumb|center|150px|Animantarx ramaljonesi"Archaeoraptor"Czerkas vide Sloan, C.P. (1999). "Feathers for T. rex?" National Geographic 196 (5 - November): pp. 98-107. vide Olson, 2000.
Hoax
Stephen Czerkas vide:C. P. Sloan vide:OlsonChimera of Yanornis and Microraptor.Atlasaurus Monbaron M. Russell D.A. Taquet P. 1999 Atlasaurus imelakeii n.g., n.sp., a brachiosaurid-like sauropod from the Middle Jurassic of Morocco Comptes Rendus de l'Académie des Sciences, Série IIA 329 7 519–526 10.1016/s1251-8050(00)80026-9 1999CRASE.329..519M gen et sp novValid
Monbaron, Russell, & TaquetMiddle JurassicTiougguit FormationMoroccoA primitive sauropod
Beipiaosaurus Xu X. Z.-L. Tang Wang X.-L 1999 A therizinosauroid dinosaur with integumentary structures from China Nature 399 6734 350–354 10.1038/20670 1999Natur.399..350X 204993327 gen et sp novValidXu, Tang, & WangEarly CretaceousYixian FormationChinaA therizinosaurthumb|center|150px|Beipiaosaurus"Bilbeyhallorum"Burge, D.L., J.H. Bird, B.K. McClelland, and M.A. Cicconetti. (1999). "Comparison of four armored dinosaurs from the Cedar Mountain Formation of Eastern Utah". Journal of Vertebrate Paleontology, Abstracts of Papers, 59th Annual Meeting, Society of Vertebrate Paleontology, Denver, CO, Volume 19, Supplement to n. 3, p. 34A.Nomen nudum
Burge, Bird, McClelland, & CicconettiAptianCedar Mountain Formation, UtahUSAinformal name for Cedarpelta.
Cedarosaurus Tidwell V. Carpenter K. Brooks W. 1999 New sauropod from the Lower Cretaceous of Utah, USA Oryctos 2 21–37 gen et sp novValid
Tidwell, Carpenter, & BrooksBarremianCedar Mountain Formation, UtahUSAA sauropod.ChaoyangsaurusZhao, Xijin Cheng, Zhengwu Xu, Xing 1999 The earliest ceratopsian from the Tuchengzi Formation of Liaoning, China Journal of Vertebrate Paleontology 19 4 681–691 10.1080/02724634.1999.10011181 1999JVPal..19..681X gen et sp novValid
Zhao, Cheng, & XuLate JurassicTuchengzi FormationChinaAn extremely primitive ceratopsianthumb|center|150px|ChaoyangsaurusDinheirosaurus Bonaparte J.F. Mateus O. 1999 A new diplodocid, Dinheirosaurus lourinhanensis gen. et sp. nov., from the Late Jurassic beds of Portugal Revista del Museo Argentino de Ciencias Naturales 5 2 13–29 gen et sp nov Valid
Bonaparte & MateusKimmeridgianAlcobaça FormationPortugalA diplodocid."Eugongbusaurus"Knoll, F. (1999). "The family Fabrosauridae". In: IV European Workshop on Vertebrate Palaeontology: p. 65;gen nov
Nomen nudum
KnollLate JurassicShishugou FormationChinaA ?hypsilophodont that hasn't yet been formally named.GondwanatitanKellner, A.W.A. and S.A.K. de Azevedo. (1999). A new sauropod dinosaur (Titanosauria) from the Late Cretaceous of Brazil. In: Proceedings of the Second Gondwanan Dinosaur Symposium (Y. Tomida, T.H. Rich, and P. Vickers-Rich, eds.): pp. 111-142.gen et sp novValid
Kellner & AzevedoLate CretaceousAdamantina FormationBrazilA titanosaurGuaibasaurusBonaparte, J.F., J. Ferigolo, and A.M. Ribeiro. (1999). A new early Late Triassic saurischian dinosaur from Rio Grande do Sul State, Brazil. In: Proceedings of the Second Gondwanan Dinosaur Symposium (Y. Tomida, T.H. Rich, and P. Vickers-Rich, eds.): pp. 89-109.gen et sp novValidBonaparte, Ferigolo, & RibeiroLate TriassicCaturrita FormationBrazilA sauropodomorph.thumb|center|150px|Guaibasaurus Jobaria Sereno P.C Allison L. Beck D.B. Dutheil H.C.E. Larsson G.H. Lyon B.M. Rudyard W. Sadleir C. Sidor C. A. Varricchio D.J. Wilson G.P. Wilson J.A. 1999 Cretaceous sauropods from the Sahara and the uneven rate of skeletal evolution among dinosaurs Science 286 5443 1342–1347 10.1126/science.286.5443.1342 10558986gen et sp novValidSereno, et alMiddle JurassicTiourarén FormationNigerA eusauropodthumb|center|150px|JobariaLessemsaurushttp://www.ameghiniana.org.ar/index.php/ameghiniana/article/view/2480 Evolución de las vértebras presacras en Sauropodomorpha Bonaparte, J. F. José Bonaparte 1999 Ameghiniana 36 115–187gen et sp novValid
BonaparteLate TriassicLos Colorados FormationArgentinaA primitive sauropodomorphthumb|center|150px|Lessemsaurus sauropoidesLirainosaurusSanz, J.L., J.E. Powell, J. Le Loueff, R. Martinez, and X. Pereda-Suberbiola. (1999). Sauropod remains from the Upper Cretaceous of Laño (Northcentral Spain). Titanosaur phylogenetic relationships. Est. Mus. Cienc. Nat. de Alava 14 (Num. Espec. 1): pp. 235-255 .gen et sp novValidSanz et alLate Cretaceous Vitoria FormationSpainA titanosaur.Lurdusaurus Taquet P. D.A. Russell 1999 A massively constructed iguandont from Gadoufaouna, Lower Cretaceous of Niger Annales de Paléontologie 85 1 85–96 10.1016/s0753-3969(99)80009-3 1999AnPal..85...85T https://hal.science/hal-04612943 gen et sp novValid
Taquet & RussellAptianElrhaz FormationNigerA hadrosaurthumb|center|150px|Lurdusaurus "Newtonsaurus"Pickering, S. (1995). A fractal scaling in dinosaurology project (2nd revised printing). Capitola, California: 478 pages; [nomen nudum].gen et sp novNomen nudumWelles vide: Welles & PickeringLate TriassicRhaetic bedsWalesInformal name. Formally describe in 2025.thumb|center|150px|"Newtonsaurus"
Nigersaurusgen et sp novValid
Sereno et alEarly CretaceousElrhaz FormationNigerA rebbachisauridthumb|center|150px|Nigersaurus Nodocephalosaurus Sullivan R.M. 1999 Nodocephalosaurus kirtlandensis, gen. et sp. nov., a new ankylosaurid dinosaur (Ornithischia: Ankylosauria) from the Upper Cretaceous Kirtland Formation (upper Campanian), San Juan Basin, New Mexico Journal of Vertebrate Paleontology 19 1 126–139 10.1080/02724634.1999.10011128 1999JVPal..19..126S gen et sp nov Valid taxonSullivanCampanianKirtland FormationUSAAn ankylosauridProyandusaurusgen novsynonym
KnollBajocianShaximiao FormationChinainformal name for Hexinlusaurus
QantassaurusRich, T. H. and P. Vickers-Rich. (1999). The Hypsilophodontidae from southeastern Australia. In: Proceedings of the Second Gondwanan Dinosaur Symposium (Y. Tomida, T.H. Rich, and P. Vickers-Rich, eds.): pp. 167-180.gen et sp novValidRich & Vickers-RichEarly CretaceousWonthaggi FormationAustraliaA possible hypsilophodontidthumb|center|150px|Qantassaurus intrepidusSantanaraptorKellner, A.W.A. (1999). "Short note on a new dinosaur (Theropoda, Coelurosauria) from the Santana Formation (Romualdo Member, Albian), Northeastern Brazil". Boletim do Museu Nacional (Rio de Janeiro, Brasil), N.S. 9: 8 pages.gen et sp novValid
KellnerCenomanianSantana FormationBrazilA theropodthumb|center|180px|Santanaraptor placidusSaturnaliaLanger, M.C., F. Abdala, M. Richter, and M.J. Benton. (1999). A sauropodomorph dinosaur from the Upper Triassic (Carnian) of southern Brazil. Compte rendu hebdomadaire des séances de l'Académie des Sciences Paris, Sciences de la Terre et des Planètes 329 : pp. 511-517.gen et sp novValid
Langer, Abdala, Richter, & BentonCarnianSanta Maria FormationBrazil
ZimbabweA basal sauropodomorphthumb|center|180px|Saturnalia
Sinornithosaurus Xu X. Wang X.-L. Wu X.-C. 1999 A dromaeosaurid dinosaur with a filamentous integument from the Yixian Formation of China Nature 401 6750 262–266 10.1038/45769 1999Natur.401..262X 4430574 gen et sp novValid
Xu, Wang, & WuAptianYixian FormationChinaA Dromaeosauridthumb|center|200px|SinornithosaurusTangvayosaurus Allain R. Taquet P. Battail B. Dejax J. Richir P. Veran M. Vacant R. Mateus O. Sayarath P. Khenthavong B. Phouyavong S. 1999 Un nouveau genre de dinosaure sauropode de la formation des Gres superieurs (Aptien-Albien) du Laos Comptes Rendus de l'Académie des Sciences, Série IIA 329 8 609–616 10.1016/s1251-8050(00)87218-3 1999CRASE.329..609A Gen et sp novValidAllain et atEarly CretaceousGrès Supérior FormationLaosA titanosaur
TehuelchesaurusRich, T.H., P. Vickers-Rich, O. Gimenez, R. Cúneo, P. Puerta, and P. Vacca. (1999). "A new sauropod dinosaur from Chubut Province, Argentina". In: Proceedings of the Second Gondwanan Dinosaur Symposium (Y. Tomida, T.H. Rich, and P. Vickers-Rich, eds.): pp. 61-84.Gen et sp nov
ValidRich et alMiddle JurassicCañadon Asfalto FormationArgentinaA eusauropod
Newly named birds
Name Novelty Status Authors Age Unit Location Notes ImagesAmplibuteo concordatus Emslie, Steven D. Czaplewski, Nicholas J. 1999 Two New Fossil Eagles from the Late Pliocene (Late Blancan) of Florida and Arizona and Their Biogeographic Implications http://www.sil.si.edu/smithsoniancontributions/paleobiology/pdf_lo/sctp-0089.pdf In: Avian Paleontology at the Close of the 20th Century: Proceedings of the 4th International Meeting of the Society of Avian Paleontology and Evolution, Washington, D.C., 4–7 June 1996, Smithsonian Contributions to Paleobiology 89 185–198 Sp. nov.
ValidSteven D. Emslie
Nicholas J. CzaplewskiLate PlioceneFlorida and ArizonaUSA:
Florida;
ArizonaAn Accipitridae.Anatalavis oxfordi Olson, Storrs L. 1999 Avian Paleontology at the Close of the 20th Century: Proceedings of the 4th International Meeting of the Society of Avian Paleontology and Evolution, Washington, D.C, 4-7 June 1996 http://www.sil.si.edu/smithsoniancontributions/paleobiology/pdf_lo/sctp-0089.pdf Smithsonian Contributions to Paleobiology 89 89 231–243 10.5479/si.00810266.89.1Sp. nov.
ValidStorrs L. OlsonEarly EoceneYpresian,
London Clay,
MP 8UK:
EnglandAn Anseranatidae, Anatalavinae Olson, 1999. In 2002 Jíří Mlíkovský made this species the type species of his not widely accepted new genus Nettapterornis.Mlíkovský, Jíří 2002 Cenozoic Birds of the World Part 1: Europe http://www.nm.cz/download/JML-18-2002-CBE.pdf Praha Ninox Press 2002 1–407 2014-08-04 https://web.archive.org/web/20110520101755/http://www.nm.cz/download/JML-18-2002-CBE.pdf 2011-05-20 dead Aquila bivia Sp. nov.
ValidSteven D. Emslie
Nicholas J. CzaplewskiLate PlioceneFloridaUSA:
Florida;
ArizonaAn Accipitridae.Argornis caucasicus Karkhu, Aleksandr A. 1999 A New Genus and Species of the Family Jungornithidae (Apodiformes) from the Late Eocene of the Northern Caucas, with Comments on the Ancestry of Hummingbirds http://www.sil.si.edu/smithsoniancontributions/paleobiology/pdf_lo/sctp-0089.pdf In: Avian Paleontology at the Close of the 20th Century: Proceedings of the 4th International Meeting of the Society of Avian Paleontology and Evolution, Washington, D.C., 4–7 June 1996, Smithsonian Contributions to Paleobiology 89 207–216 Gen. nov. et Sp. nov.
ValidAleksandr A. KarkhuLate EoceneNorthern CaucasusRussiaAn Apodiformes, Jungornithidae Karkhu, 1988.Canadaga arctica Hou Lianhai 1999 New Hesperornithid (Aves) from the Canadian Arctic http://www.ivpp.cas.cn/cbw/gjzdwxb/xbwzxz/200812/W020090813369411425606.pdf Vertebrata PalAsiatica 37 3 228–233 10.19615/j.cnki.1000-3118.1999.03.009 Gen. nov. et Sp. nov.
ValidHou LianhaiLate CretaceousMiddle MaastrichtianCanada:
NunavutA Hesperornithidae Marsh, 1872.Changchengornis hengdaoziensis Qiang, Ji Chiappe, Luis M. Shu’an, Ji 1999 A New Late Mesozoic Confuciusornithid Bird from China Journal of Vertebrate Paleontology 19 1 1–7 10.1080/02724634.1999.10011117 1999JVPal..19....1Q Gen. nov. et Sp. nov.
ValidJi Qiang
Luis M. Chiappe
Ji Shu’anEarly CretaceousChaomidianzi FormationChinaA Confuciusornithidae Hou, Zhou, Gu et Zhang, 1995.Chaunoides antiquus de Alvarenga, Herculano M. F. 1999 A Fossil Screamer (Anseriformes: Anhimidae) from the Middle Tertiary of Southeastern Brazil http://www.sil.si.edu/smithsoniancontributions/paleobiology/pdf_lo/sctp-0089.pdf In: Avian Paleontology at the Close of the 20th Century: Proceedings of the 4th International Meeting of the Society of Avian Paleontology and Evolution, Washington, D.C., 4–7 June 1996, Smithsonian Contributions to Paleobiology 89 223–230 Sp. nov.
ValidHerculano M. F. de AlvarengaLate Oligocene or Early MioceneTremembé FormationBrazilAn Anhimidae.Chauvireria bulgarica Boev, Zlatozar N. 1999 Neogene-Quaternary Birds of Bulgaria https://www.researchgate.net/publication/301226810 София 508 Sp. nov.
ValidBoevPleistoceneBulgariaA Phasianidae.Confuciusornis dui Hou, Lianhai Martin, Larry D. Zhou, Zhonghe Feduccia, Alan Zhang, Fucheng 1999 A Diapsid Skull in a New Species of the Primitive Bird Confuciusornis Nature 399 6737 679–682 10.1038/21411 1999Natur.399..679H 4402195 Sp. nov.
ValidHou Lianhai
Larry D. Martin
Zhou Zhonghe
Alan Feduccia
Zhang FuchengEarly CretaceousYixian FormationChinaA Confuciusornithidae Hou, Zhou, Gu et Zhang, 1995.Dryolimnas augusti Mourer-Chauviré, Cécile Bour, Roger Ribes, Sonia Moutou, François 1999 The Avifauna of Réunion Island (Mascarene Islands) at the Time of the Arrival of the First Europeans http://www.sil.si.edu/smithsoniancontributions/paleobiology/pdf_lo/sctp-0089.pdf In: Avian Paleontology at the Close of the 20th Century: Proceedings of the 4th International Meeting of the Society of Avian Paleontology and Evolution, Washington, D.C., 4–7 June 1996, Smithsonian Contributions to Paleobiology 89 1–38 Sp. nov.
ValidCécile Mourer-Chauviré
Roger Bour
Sonia Ribes
François MoutouHoloceneRéunionRéunionA Rallidae.Emberiza alcoveri Rando, J. C. Lopez, M. Seguí, B. 1999 A New Species of Extinct Flightless Passerine (Emberizidae: Emberiza) from the Canary Islands http://sora.unm.edu/sites/default/files/journals/condor/v101n01/p0001-p0013.pdf The Condor 101 1 1–13 10.2307/1370440 1370440 Sp. nov.
ValidJ. C. Rando
M. Lopez
B. SeguíLate Pleistocene-HoloceneCanary Islands, TenerifeSpain:
Canary IslandsAn Emberizidae.Eocuculus cherpinae Chandler, Robert M. 1999 Fossil Birds of Florissant, Colorado: With a Description of a New Genus and Species of Cuckoo The National Park Service Paleontological Research 4 49–53 Gen. nov. et Sp. nov.
ValidRobert M. ChandlerLate EoceneEarly ChadronianUSA:
ColoradoA ?Cuculidae.Eoenantiornis buhleri Hou, Lianhai Martin, Larry D. Zhou, Zongzhe Feduccia, Alan 1999 Archaeopteryx to Opposite Birds Missing Link from the Mesozoic of China http://www.ivpp.cas.cn/cbw/gjzdwxb/xbwzxz/200812/W020090813369438394257.pdf Vertebrata PalAsiatica 37 2 88–95 10.19615/j.cnki.1000-3118.1999.02.002 Gen. nov. et Sp. nov.
ValidHou Lianhai
Larry D. Martin
Zhou Zongzhe
Alan FeducciaEarly CretaceousYixian FormationChinaAn Enantiornithes Walker, 1981, Eoenantiornithiformes Hou, Martin, Zhou et Feduccia, 1999, Eoenantiornithidae Hou, Martin, Zhou et Feduccia, 1999, this is the type species of the new genus.Falco bakalovi Boev, Zlatozar N. 1999 Falco bakalovi sp. n. - a Late Pliocene Falcon (Falconidae, Aves) from Varhets (W Bulgaria) Geologica Balcanica 29 1–2 131–135 10.52321/GeolBalc.29.1-2.131 255926847 free Sp. nov.
ValidZlatozar N. BoevLate PlioceneVillafranchian,
MN 17BulgariaA Falconidae.Gallinula balcanica Boev, Zlatozar N. 1999 Gallinula balcanica sp. n. - a Villafranchian Moorhen from Bulgaria Acta Zoologica Bulgarica 51 43–47 Sp. nov.
ValidZlatozar N. BoevLate PlioceneVillafranchian,
MN 17BulgariaA Rallidae.Graculavus augustus Hope, Sylvia 1999 A New Species of Graculavus from the Cretaceous of Wyoming (Aves: Neornithes) http://www.sil.si.edu/smithsoniancontributions/paleobiology/pdf_lo/sctp-0089.pdf In: Avian Paleontology at the Close of the 20th Century: Proceedings of the 4th International Meeting of the Society of Avian Paleontology and Evolution, Washington, D.C., 4–7 June 1996, Smithsonian Contributions to Paleobiology 89 261–266 Sp. nov.
ValidSylvia HopeLate CretaceousLate Maastrichtian,
Lance FormationUSA:
WyomingA Graculavidae Fürbringer, 1888.Gurilynia nessovi Kurochkin, Evgeny N. 1999 A New Large Enantiornithid from the Upper Cretaceous of Mongolia (Aves, Enantiornithes) Russian Academy of Sciences, Proceedings of the Zoological Institute 277 130–141 Gen. nov. et Sp. nov.
ValidEvgeny N. KurochkinLate CretaceousGurilyn TsavMongoliaAn Enantiornithes Walker, 1981, Alexornithiformes Brodkorb, 1976, Enantiornithidae Nessov & Borkin, 1983.Hebeiornis fengningensis Yan in Xu Gui-lin, Yang You-shi & Deng Shao-yeng 1999 First Discovery of Mesozoic Bird Fossils in Hebei Province and Its Significance Regional Geology of China 18 444–448 Gen. nov. et Sp. nov.
ValidYanEarly CretaceousYixian FormationChinaOrnithothoraces Chiappe et Calvo, 1994, this is the type species of the new genus. Vescornis hebeiensis Zhang, Ericson et Zhou, 2004. is a junior synonym.Zhang, Fucheng Ericson, Per G.P. Zhou, Zhonghe 2004 Description of a New Enantiornithine Bird from the Early Cretaceous of Hebei, Northern China http://www.ivpp.ac.cn/qt/papers/201206/P020120604365639534651.pdf Canadian Journal of Earth Sciences 41 9 1097–1107 10.1139/e04-055 2004CaJES..41.1097Z 2014-08-04 https://web.archive.org/web/20141023131246/http://www.ivpp.ac.cn/qt/papers/201206/P020120604365639534651.pdf 2014-10-23 dead Liaoxiornis delicatus Hou Lianhai Chen Peiji 1999 Liaoxiornis delicatus gen. et sp. nov., the Smallest Mesozoic Bird http://csb.scichina.com/kxtbe/EN/abstract/abstract313063.shtml# Chinese Science Bulletin 44 9 834–838 10.1007/bf02885031 dead https://web.archive.org/web/20140808050232/http://csb.scichina.com:8080/kxtbe/EN/abstract/abstract313063.shtml 2014-08-08 1999ChSBu..44..834H 84048214 subscription Gen. nov. et Sp. nov.
ValidHou Lianhai
Chen PeijiEarly CretaceousYixian FormationChinaA Liaoxiornithidae L. H. Hou et Chen, 1999, this is the type species of the new genus. Lingyuanornis parvus Ji & Ji, 1999. is a junior synonym.Ji Qiang Ji Shuan 1999 A New Genus of Mesozoic Birds from Lingyuan, Liaoning, China http://en.cnki.com.cn/Article_en/CJFDTOTAL-DIZI199903021.htm Chinese Geology, 1999 262 3 45–48 Longmornis robustirostrata Boles, Walter E. 1999 A Songbird (Aves: Passeriformes: Oriolidae) from the Miocene of Riversleigh, Northwestern Queensland, Australia https://www.researchgate.net/publication/232943138 Alcheringa 23 1 51–56 10.1080/03115519908619338 1999Alch...23...51B Gen, nov. et Sp. nov.
ValidWalter E. Boles?Early MioceneRiversleighAustralia:
QueenslandAn Oriolidae, this is the type species of the new genus.Loxia patevi Boev, Zlatozar N. 1999 Earliest Finds of Crossbills (Genus Loxia) (Aves: Fringillidae) from Varhshets (NW Bulgaria) Geologica Balcanica 29 3–4 51–57 10.52321/GeolBalc.29.3-4.51 255877416 free Sp. nov.
ValidZlatozar N. BoevLate PlioceneMN 17BulgariaA Fringillidae.Masillapodargus longipesMayr, Gerald 1999 Caprimulgiform Birds from the Middle Eocene of Messel (Hessen, Germany) Journal of Vertebrate Paleontology 79 3 521–532 10.1080/02724634.1999.10011162 1999JVPal..19..521M Gen. nov. et Sp. nov.
ValidGerald MayrMiddle EoceneMessel pit,
MP 11Germany:
HessenA stem Podargidae, this is the type species of the new genus.Ngawupodius minya Boles, Walter E. Ivison, Tessa J. 1999 A New Genus of Dwarf Megapode (Galliformes: Megapodiidae) from the Late Oligocene of Central Australia http://www.sil.si.edu/smithsoniancontributions/paleobiology/pdf_lo/sctp-0089.pdf In: Avian Paleontology at the Close of the 20th Century: Proceedings of the 4th International Meeting of the Society of Avian Paleontology and Evolution, Washington, D.C., 4–7 June 1996, Smithsonian Contributions to Paleobiology 89 199–206 Gen. nov. et Sp. nov.ValidWalter E. Boles
Tessa J. IvisonLate OligoceneNamba FormationAustralia:
South AustraliaA stem Megapodiidae, this is the type species of the new genus.Nupharanassa bohemica Mlíkovský, Jirí 1999 A New Jacana (Aves: Jacanidae) from the Early Miocene of the Czech Republic http://www.nm.cz/download/pm/zoo/mlikovsky_lit/130-1999-MioceneJacanidae.pdf Comptes Rendus de l'Académie des Sciences, Série II 328 2 121–123 10.1016/s1251-8050(99)80007-x 1999CRASE.328..121M 2014-08-04 https://web.archive.org/web/20140808053630/http://www.nm.cz/download/pm/zoo/mlikovsky_lit/130-1999-MioceneJacanidae.pdf 2014-08-08 dead Sp. nov.Valid ?Jirí MlíkovskýEarly MioceneMN 4bCzech RepublicA Jacanidae. Mourer-Chauviré, 1999 transferred this species to the genus Geranopterus Milne-Edwards, 1892 and later it was placed in the Geranopteridae Mayr & Mourer-Chauviré, 2000.Mourer-Chauviré, Cécile 1999 Position Systématique de Nupharanassa bohemica Mlikovsky, 1999 https://www.researchgate.net/publication/251663116 329 2 149–152 10.1016/s1251-8050(99)80217-1 Comptes Rendus de l'Académie des Sciences, Série IIA 1999CRASE.329..149M Paraprefica kelleriGen. nov. et Sp. nov.ValidGerald MayrMiddle EoceneMessel pit,
MP 11Germany:
HessenA stem Nyctibiidae, this is the type species of the new genus.Paraprefica majorSp. nov.ValidGerald MayrMiddle EoceneMessel pit,
MP 11Germany:
HessenA stem Nyctibiidae.Pelecanus schreiberi Olson, Strorrs L. 1999 A New Species of Pelican (Aves: Pelecanidae) from the Lower Pliocene of North Carolina and Florida http://biostor.org/cache/pdf/90/84/e6/9084e6c696b8c655ce8fe5584c2acb61.pdf Proceedings of the Biological Society of Washington 112 3 503–509 2014-08-04 https://web.archive.org/web/20140808060637/http://biostor.org/cache/pdf/90/84/e6/9084e6c696b8c655ce8fe5584c2acb61.pdf 2014-08-08 dead Sp. nov.ValidStrorrs L. OlsonEarly PlioceneYorktown FormationUSA:
North CarolinaA Pelecanidae.Primotrogon wintersteiniMayr, Gerald 1999 A New Trogon from the Middle Oligocene of Céreste, France http://www.senckenberg.de/files/content/forschung/abteilung/terrzool/ornithologie/primotrogon.pdf The Auk 116 2 427–434 10.2307/4089376 4089376 Gen. nov. et Sp. nov.ValidGerald MayrMiddle OligoceneMP 23FranceA stem Trogonidae, the type species of the new genus.Pumiliornis tessellatusMayr, Gerald 1999 Pumiliornis tessellatus n. gen. n. sp., A New Enigmatic Bird from the Middle Eocene of Grube Messel (Hessen, Germany) http://www.senckenberg.de/files/content/forschung/abteilung/terrzool/ornithologie/pumiliornis.pdf Courier Forschungsinstitut Senckenberg 216 75–83 Gen. nov. et Sp. nov.ValidGerald MayrMiddle EoceneMessel pit,
MP 11Germany:
HessenOrder and family Incerta Sedis, the type species of the new genus.Regulus bulgaricus Boev, Zlatozar N. 1999 Regulus bulgaricus sp. n. - The First Fossil Kinglet (Aves: Sylviidae) from the Late Pliocene of Varshets, Western Bulgaria http://biostor.org/cache/pdf/2f/ad/a3/2fada3063ba8f07da48871a8365d5868.pdf Historia Naturalis Bulgarica 10 109–115 dead https://web.archive.org/web/20140720112439/http://biostor.org/cache/pdf/2f/ad/a3/2fada3063ba8f07da48871a8365d5868.pdf 2014-07-20 Sp. nov.ValidZlatozar N. BoevLate PlioceneMN 17BulgariaA Regulidae.Scolopax carmesinae Seguí, Bartomeu 1999 A Late Tertiary Woodcock from Menorca, Balearic Islands, Western Mediterranean https://sora.unm.edu/sites/default/files/journals/condor/v101n04/p0909-p0915.pdf The Condor 101 4 909–915 10.2307/1370088 1370088 Sp. nov.ValidBartomeu SeguíLate PlioceneMenorca,
MN 17-18Spain:
MenorcaA Scolopacidae.Selmes absurdipes Peters, Dieter S. 1999 Selmes absurdipes, New Genus, New Species, a Sancoleiform Bird from the Oil Shale of Messel (Germany, Middle Eocene) http://www.sil.si.edu/smithsoniancontributions/paleobiology/pdf_lo/sctp-0089.pdf In: Avian Paleontology at the Close of the 20th Century: Proceedings of the 4th International Meeting of the Society of Avian Paleontology and Evolution, Washington, D.C., 4–7 June 1996, Smithsonian Contributions to Paleobiology 89 217–222 Gen. nov. et Sp. nov.ValidDieter S. PetersMiddle EoceneMessel pit,
MP 11;
MP 13Germany:
Hessen;
Saxony-AnhaltA Coliiformes, Selmeidae Zelenkov et Dyke, 2008, the type species of the new genus.Tyto neddi Steadman, David W. Hilgartner, William B. 1999 A New Species of Extinct Barn Owl (Aves: Tyto) from Barbuda, Lesser Antilles http://www.sil.si.edu/smithsoniancontributions/paleobiology/pdf_lo/sctp-0089.pdf In: Avian Paleontology at the Close of the 20th Century: Proceedings of the 4th International Meeting of the Society of Avian Paleontology and Evolution, Washington, D.C., 4–7 June 1996, Smithsonian Contributions to Paleobiology 89 75–83 Sp. nov.ValidDavid W. Steadman
William B. HilgartnerLate Pleistocene-Early HoloceneBarbudaAntigua and BarbudaA Tytonidae.Yandangornis longicaudus Cai Zhengquan Zhao Lijun 1999 A Long Tailed Bird from the Late Cretaceous of Zhejiang Science in China Series D: Earth Sciences 42 4 434–441 http://earth.scichina.com:8080/sciDe/EN/abstract/abstract306212.shtml# 2014-08-03 https://web.archive.org/web/20160610010623/http://earth.scichina.com:8080/sciDe/EN/abstract/abstract306212.shtml 2016-06-10 dead Gen. nov. et Sp. nov.ValidCai Zhengquan
Zhao LijunEarly Late CretaceousSantonianChinaAn Enentiornithes, this is the type species of the new genus.
Newly named pterosaurs
Name Novelty Status Authors Age Unit Location Notes Images
Dendrorhynchoides Ji S.-A. Ji Q. Padian K. 1999 Biostratigraphy of new pterosaurs from China Nature 398 6728 573–574 10.1038/19221 1999Natur.398..573J 4316586 free gen nov
Valid
Ji S.-A., Ji Q., & PadianAptianYixian FormationChinaA new genus for "Dendrorhynchus" curvidentatus (Ji & Ji, 1998)
Siroccopteryxgen et sp nov
junior synonym?
Mader & KellnerCenomanianKem Kem FormationMoroccopossibly a jr synonym of Coloborhynchus?
TendaguripterusUnwin, David M. Heinrich, Wolf-Dieter 1999 On a pterosaur jaw from the Upper Jurassic of Tendaguru (Tanzania) Mitteilungen aus dem Museum für Naturkunde in Berlin, Geowissenschaftliche Reihe 2 1 121–134 10.1002/mmng.1999.4860020109 1999FossR...2..121U free gen et sp nov
Valid
Unwin & HeinrichKimmeridgian
Tendaguru FormationTanzaniaMonotypic with the species T. recki
Lepidosauromorphs
Newly named plesiosaurs
Name Novelty Status Authors Age Unit Location Notes Images
Callawayasaurus Carpenter K 1999 Revision of North American elasmosaurs from the Cretaceous of the Western Interior Paludicola 2 2 148–173 gen nov
ValidCarpenterAptianPaja FormationColombia
New genus for the species "Alzadasaurus" colombiensis Wells, 1962
Newly named scincomorphans
Name Novelty Status Authors Age Unit Location Notes ImagesPachygenys Gao K. Cheng Z. 1999 A new lizard from the Lower Cretaceous of Shandong, China Journal of Vertebrate Paleontology 19 3 456–465 4524009 10.1080/02724634.1999.10011158 1999JVPal..19..456K gen nov
ValidGao and ChengEarly CretaceousEastern ChinaChina
Has a distinctly small amount of dentary teeth.
Other Animals
Name Status Authors Age Unit Location Notes Images Charniodiscus yorgensis Disputed Borchvardt et Nessov Ediacaran Zimnie Gory Formation and Ust-Pinega Formation Russia
References
30em
Category:1990s in paleontology
Paleontology