2014 in reptile paleontology

none 2014 reptile paleontology science paleobotany paleontology arthropod paleontology paleoentomology paleomalacology archosaur paleontology mammal paleontology paleoichthyology Paleontology History of science Dinosaurs This list of fossil reptiles described in 2014 is a list of new taxa of fossil reptiles that were described during the year 2014, as well as other significant discoveries and events related to reptile paleontology that occurred in 2014. Ichthyopterygians Research Anatomy, taxonomy and phylogenetic relationships of Cretaceous ophthalmosaurids Simbirskiasaurus birjukovi and Pervushovisaurus bannovkensis are reevaluated by Fischer et al. (2014).Valentin Fischer Maxim S. Arkhangelsky Darren Naish Ilya M. Stenshin Gleb N. Uspensky Pascal Godefroit amp 2014 Simbirskiasaurus and Pervushovisaurus reassessed: implications for the taxonomy and cranial osteology of Cretaceous platypterygiine ichthyosaurs Zoological Journal of the Linnean Society 171 4 822–841 10.1111/zoj.12158 67765108 http://www.vliz.be/imisdocs/publications/346572.pdf New taxa Name Novelty Status Authors Age Unit Location Notes ImagesJanusaurusAubrey Jane Roberts Patrick Scott Druckenmiller Glenn-Peter Sætre Jørn Harald Hurum amp 2014 A New Upper Jurassic Ophthalmosaurid Ichthyosaur from the Slottsmøya Member, Agardhfjellet Formation of Central Spitsbergen PLOS ONE 9 8 e103152 10.1371/journal.pone.0103152 25084533 4118863 2014PLoSO...9j3152R free Gen. et sp. novValidRoberts et al.Late JurassicAgardhfjellet FormationNorwayAn ophthalmosaurid ichthyosaur. The type species is Janusaurus lundi. Zverkov & Prilepskaya (2019) considered Janusaurus to be a junior synonym of the genus Arthropterygius, though the authors maintained J. lundi as a distinct species within the latter genus;Nikolay G. Zverkov Natalya E. Prilepskaya 2019 A prevalence of Arthropterygius (Ichthyosauria: Ophthalmosauridae) in the Late Jurassic—earliest Cretaceous of the Boreal Realm PeerJ 7 e6799 10.7717/peerj.6799 31106052 6497043 free Delsett et al. (2019) rejected this synonymy.Lene L. Delsett Aubrey J. Roberts Patrick S. Druckenmiller Jørn H. Hurum 2019 Osteology and phylogeny of Late Jurassic ichthyosaurs from the Slottsmøya Member Lagerstätte (Spitsbergen, Svalbard) Acta Palaeontologica Polonica 64 4 717–743 10.4202/app.00571.2018 210294877 free 10852/76089 free LeniniaValentin Fischer Maxim S. Arkhangelsky Gleb N. Uspensky Ilya M. Stenshin Pascal Godefroit Pascal Godefroit amp 2014 A new Lower Cretaceous ichthyosaur from Russia reveals skull shape conservatism within Ophthalmosaurinae Geological Magazine 151 1 60–70 10.1017/S0016756812000994 2014GeoM..151...60F 2268/137853 49470362 http://osf.io/t3j7q/ free Gen. et sp. novValidFischer et al.Early Cretaceous (early Aptian)RussiaAn ophthalmosaurine ophthalmosaurid ichthyosaur. The type species is Leninia stellans.SisteroniaValentin Fischer Nathalie Bardet Myette Guiomar Pascal Godefroit amp 2014 High Diversity in Cretaceous Ichthyosaurs from Europe Prior to Their Extinction PLOS ONE 9 1 e84709 10.1371/journal.pone.0084709 24465427 3897400 2014PLoSO...984709F free Gen. et sp. novValidFischer et al.Cretaceous (early Albian to early Cenomanian)Cambridge Greensand Gault Formation Marnes Bleues FormationFrance United KingdomA platypterygiine ophthalmosaurid ichthyosaur. The type species is Sisteronia seeleyi.200px|Undorosaurus trautscholdiM.S. Arkhangelsky N.G. Zverkov amp 2014 On a new ichthyosaur of the genus Undorosaurus Proceedings of the Zoological Institute of the Russian Academy of Sciences 318 3 187–196 http://www.zin.ru/journals/trudyzin/doc/vol_318_3/TZ_318_3_Arkhangelsky.pdf Sp. novValidArkhangelsky & ZverkovLate JurassicRussiaAn ophthalmosaurine ophthalmosaurid ichthyosaur, a species of Undorosaurus. Sauropterygians Name Novelty Status Authors Age Unit Location Notes ImagesAristonectes quiriquinensisRodrigo A. Otero Sergio Soto-Acuña Frank Robin O'Keefe José P. O'Gorman Wolfgang Stinnesbeck Mario E. Suárez David Rubilar-Rogers Christian Salazar Luis Arturo Quinzio-Sinn 2014 Aristonectes quiriquinensis, sp. nov., a new highly derived elasmosaurid from the upper Maastrichtian of central Chile Journal of Vertebrate Paleontology 34 1 100–125 10.1080/02724634.2013.780953 11336/79718 84729992 http://sedici.unlp.edu.ar/handle/10915/128448 free Sp. nov ValidOtero et al.Late Cretaceous (late Maastrichtian)Quiriquina FormationChileA plesiosaur, a species of Aristonectes.200px|Cymatosaurus erikaeMichael W. Maisch 2014 A well preserved skull of Cymatosaurus (Reptilia: Sauropterygia) from the uppermost Buntsandstein (Middle Triassic) of Germany Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen 272 2 213–224 10.1127/0077-7749/2014/0407 Sp. nov ValidMaischMiddle Triassic (earliest Anisian)Röt FormationGermanyA species of Cymatosaurus.MajiashanosaurusDa-Yong Jiang Ryosuke Motani Andrea Tintori Olivier Rieppel Guan-Bao Chen Jian-Dong Huang Rong Zhang Zuo-Yu Sun Cheng Ji 2014 The Early Triassic eosauropterygian Majiashanosaurus discocoracoidis, gen. et sp. nov. (Reptilia, Sauropterygia), from Chaohu, Anhui Province, People's Republic of China Journal of Vertebrate Paleontology 34 5 1044–1052 10.1080/02724634.2014.846264 129828376 Gen. et sp. nov ValidJiang et al.Early Triassic (Olenekian)Nanlinghu FormationChinaA basal member of Eosauropterygia. The type species is Majiashanosaurus discocoracoidis.200px|Nothosaurus zhangiJun Liu Shi-xue Hu Olivier Rieppel Da-yong Jiang Michael J. Benton Neil P. Kelley Jonathan C. Aitchison Chang-yong Zhou Wen Wen Jin-yuan Huang Tao Xie Tao Lv 2014 A gigantic nothosaur (Reptilia: Sauropterygia) from the Middle Triassic of SW China and its implication for the Triassic biotic recovery Scientific Reports 4 7142 10.1038/srep07142 free 25429609 4245812 Sp. novValidLiu et al.Middle Triassic (Anisian)Guanling FormationChinaA nothosaur, a species of Nothosaurus.OdoiporosaurusSilvio Renesto Giorgio Binelli Hans Hagdorn 2014 A new pachypleurosaur from the Middle Triassic Besano Formation of Northern Italy Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen 271 2 151–168 10.1127/0077-7749/2014/0382 Gen. et sp. nov ValidRenesto, Binelli & HagdornMiddle Triassic (Anisian)Besano FormationItalyA pachypleurosaur. The type species is Odoiporosaurus teruzzii.200px|PararcusNicole Klein Torsten M. Scheyer amp 2014 A new placodont sauropterygian from the Middle Triassic of the Netherlands Acta Palaeontologica Polonica 59 4 887–902 10.4202/app.2012.0147 54714765 free Gen. et sp. novValidKlein & ScheyerMiddle Triassic (early Anisian)Vossenveld FormationNetherlandsA placodont. The type species is Pararcus diepenbroeki.200px|Pliosaurus patagonicusZulma Gasparini José P. O'Gorman 2014 A new species of Pliosaurus (Sauropterygia, Plesiosauria) from the Upper Jurassic of northwestern Patagonia, Argentina Ameghiniana 51 4 269–283 http://www.ameghiniana.org.ar/index.php/ameghiniana/article/view/2225 10.5710/amgh.03.04.2014.2225 130194647 11336/9372 free Sp. nov ValidGasparini & O'GormanLate Jurassic (middle Tithonian)Vaca Muerta FormationArgentinaA species of Pliosaurus. Lepidosaurs Newly named rhynchocephalians Name Novelty Status Authors Age Unit Location Notes ImagesKawasphenodon peligrensisSebastian Apesteguia Raúl O. Gómez Guillermo W. Rougier 2014 The youngest South American rhynchocephalian, a survivor of the K/Pg extinction Proceedings of the Royal Society B: Biological Sciences 281 1792 20140811 10.1098/rspb.2014.0811 25143041 4150314Sp. novValidApesteguia, Gómez & RougierPalaeoceneSalamanca FormationArgentinaAn opisthodontian sphenodontid, a species of Kawasphenodon.Priosphenodon minimusSebastian Apesteguia Jose L. Carballido 2014 A new eilenodontine (Lepidosauria, Sphenodontidae) from the Lower Cretaceous of central Patagonia Journal of Vertebrate Paleontology 34 2 303–317 10.1080/02724634.2013.803974 11336/6524 130381621 free Sp. novValidApesteguia & CarballidoEarly Cretaceous Cerro Barcino FormationArgentinaAn eilenodontine sphenodontid, a species of Priosphenodon. Newly named lizards Name Novelty Status Authors Age Unit Location Notes ImagesBlanus mendeziArnau Bolet Massimo Delfino Josep Fortuny Sergio Almécija Josep M. Robles David M. Alba 2014 An Amphisbaenian Skull from the European Miocene and the Evolution of Mediterranean Worm Lizards PLOS ONE 9 6 e98082 10.1371/journal.pone.0098082 24896828 4045672 2014PLoSO...998082B free Sp. novValidBolet et al.Miocene (11.6 Mya)Vallès-Penedès BasinSpainAn amphisbaenian, a species of Blanus.Blanus thomaskelleriAndrej Čerňanský Jean-Claude Rage Jozef Klembara 2014 The Early Miocene squamates of Amöneburg (Germany): the first stages of modern squamates in Europe Journal of Systematic Palaeontology 13 2 97–128 10.1080/14772019.2014.897266 140170242 Sp. novValidČerňanský, Rage & KlembaraEarly MioceneGermanyAn amphisbaenian, a species of Blanus.CalangubanTiago R. Simões Michael W. Caldwell Alexander W. A. Kellner 2014 A new Early Cretaceous lizard species from Brazil, and the phylogenetic position of the oldest known South American squamates Journal of Systematic Palaeontology 13 7 601–614 10.1080/14772019.2014.947342 84446189 Gen. et sp. novValidSimões, Caldwell & KellnerEarly CretaceousCrato FormationBrazilOriginally described as a scleroglossan lizard with scincomorph affinities; subsequently argued to be a borioteiioid.Ednalva da Silva Santos Samuel Cardozo Ribeiro Antônio Álamo Feitosa Saraiva Alexander Wilhelm Armin Kellner Tiago R. Simões A neotype and reassessment of phylogenetic relationships of the fossil lizard Calanguban alamoi from the Crato Formation (Early Cretaceous, Brazil) Journal of Systematic Palaeontology 2025 23 1 2496536 10.1080/14772019.2025.2496536 The type species is Calanguban alamoi.DakotasepsRandall L. Nydam 2014 Dakotaseps gen. nov., a replacement name for the lizard genus Dakotasaurus Nydam 2013, a junior homonym of the ichnotaxon Dakotasaurus Branson and Mehl 1932 Zootaxa 3900 1 150 10.11646/zootaxa.3900.1.11 25543731 free Nom. nov ValidNydamLate Cretaceous (Cenomanian)Dakota FormationUnited StatesA member of Scincomorpha of uncertain phylogenetic placement, assigned by Nydam (2013) to an informal paramacellodid-cordylid grade;Randall L. Nydam 2013 Lizards and Snakes from the Cenomanian through Campanian of Southern Utah: Filling the Gap in the Fossil Record of Squamata from the Late Cretaceous of the Western Interior of North America Alan L. Titus Mark A. Loewen At the Top of the Grand Staircase: The Late Cretaceous of Southern Utah Indiana University Press 370–423 978-0-253-00896-1 a replacement name for Dakotasaurus Nydam (2013) (preoccupied).FuniusaurusLi Xu Xiaochun Wu Junchang Lü Songhai Jia Jiming Zhang Hanyong Pu Xingliao Zhang 2014 A New Lizard (Lepidosauria: Squamata) from the Upper Cretaceous of Henan, China Acta Geologica Sinica - English Edition 88 4 1041–1050 10.1111/1755-6724.12271 128552980 Gen. et sp. novValidXu et al.Late CretaceousQiupa FormationChinaA member of Polyglyphanodontidae. The type species is Funiusaurus luanchuanensis.MaioricalacertaSalvador Bailon Renaud Boistel Pere Bover Josep Antoni Alcover 2014 Maioricalacerta rafelinensis, gen. et sp. nov. (Squamata, Lacertidae), from the early Pliocene of Mallorca (Balearic Islands, western Mediterranean Sea) Journal of Vertebrate Paleontology 34 2 318–326 10.1080/02724634.2013.799481 49559770 Gen. et sp. novValidBailon et al.Early Pliocene (Zanclean)SpainA member of Lacertidae. The type species is Maioricalacerta rafelinensis.PyrenasaurusArnau Bolet Marc Augé 2014 A New Miniaturized Lizard From the Late Eocene of France and Spain The Anatomical Record 297 3 505–515 10.1002/ar.22855 24482323 205411659 free Gen. et sp. novValidBolet & AugéLate EoceneFrance SpainA member of Scincoidea, possibly a skink. The type species is Pyrenasaurus evansae. Newly named snakes Name Novelty Status Authors Age Unit Location Notes ImagesAdinophisAdam C. Pritchard Jacob A. McCartney David W. Krause Nathan J. Kley 2014 New snakes from the Upper Cretaceous (Maastrichtian) Maevarano Formation, Mahajanga Basin, Madagascar Journal of Vertebrate Paleontology 34 5 1080–1093 10.1080/02724634.2014.841706 84867457 Gen. et sp. novValidPritchard et al.Late Cretaceous (Maastrichtian)Maevarano FormationMadagascarA member of Madtsoiidae. The type species is Adinophis fisaka.Indophis fanambinanaSp. novValidPritchard et al.Late Cretaceous (Maastrichtian)Maevarano FormationMadagascarA member of Nigerophiidae, a species of Indophis.RukwanyokaJacob A. McCartney Nancy J. Stevens Patrick M. O'Connor 2014 The Earliest Colubroid-Dominated Snake Fauna from Africa: Perspectives from the Late Oligocene Nsungwe Formation of Southwestern Tanzania PLOS ONE 9 3 e90415 10.1371/journal.pone.0090415 24646522 3960104 2014PLoSO...990415M free Gen. et sp. novValidMcCartney et al.OligoceneNsungwe FormationTanzaniaThe most ancient booid snake and boa from Africa. The type species is Rukwanyoka holmani.SeismophisAnnie S. Hsiou Adriana M. Albino Manuel A. Medeiros Ronny A.B. Santos 2014 The oldest Brazilian snakes from the early Late Cretaceous (Cenomanian) Acta Palaeontologica Polonica 59 3 635–642 10.4202/app.2012.0091 55266958 free 11336/99860 free Gen. et sp. novValidHsiou et al.Late Cretaceous (Cenomanian)Alcântara FormationBrazilA snake of uncertain phylogenetic placement, possibly a relative of Najash rionegrina. The type species is Seismophis septentrionalis. Turtles Research Redescription of the rediscovered holotype of the Jurassic turtle Plesiochelys etalloni is published by Anquetin, Deschamps & Claude (2014).Jérémy Anquetin Sylvie Deschamps Julien Claude 2014 The rediscovery and redescription of the holotype of the Late Jurassic turtle Plesiochelys etalloni PeerJ 2 e258 10.7717/peerj.258 24688842 3932733 free Newly named turtles Name Novelty Status Authors Age Unit Location Notes ImagesAlienochelysFrance de Lapparent de Broin Nathalie Bardet Mbarek Amaghzaz Saïd Meslouh 2014 A strange new chelonioid turtle from the Latest Cretaceous Phosphates of Morocco Comptes Rendus Palevol 13 2 87–95 10.1016/j.crpv.2013.07.008 Gen. et sp. novValidde Lapparent de Broin et al.Late Cretaceous (Maastrichtian)Oulad Abdoun BasinMoroccoA sea turtle, a member of Dermochelyoidae. The type species is Alienochelys selloumi.200px|Allaeochelys libycaPhilipe E. Havlik Walter G. Joyce Madelaine Böhme 2014 Allaeochelys libyca, a New Carettochelyine Turtle from the Middle Miocene (Langhian) of Libya Bulletin of the Peabody Museum of Natural History 55 2 201–214 10.3374/014.055.0207 56023005 http://doc.rero.ch/record/233166/files/joy_aln.pdf Sp. novValidHavlik, Joyce & BöhmeMiocene (Langhian)LibyaA relative of the pig-nosed turtle, a species of the (possibly paraphyletic) genus Allaeochelys.AshleychelysRobert E. Weems Albert E. Sanders 2014 Oligocene pancheloniid sea turtles from the vicinity of Charleston, South Carolina, U.S.A. Journal of Vertebrate Paleontology 34 1 80–99 10.1080/02724634.2013.792826 128543145 https://figshare.com/articles/journal_contribution/963533 Gen. et sp. nov ValidWeems & SandersOligoceneUnited StatesA pancheloniid sea turtle. The type species is Ashleychelys palmeri.AtolchelysPedro S. R. Romano Valéria Gallo Renato R. C. Ramos Luzia Antonioli 2014 Atolchelys lepida, a new side-necked turtle from the Early Cretaceous of Brazil and the age of crown Pleurodira Biology Letters 10 7 20140290 10.1098/rsbl.2014.0290 25079494 4126622Gen. et sp. novValidRomano et al.Early Cretaceous (Barremian)Morro do Chaves FormationBrazilA bothremydid pleurodiran. The type species is Atolchelys lepida.Brodiechelys royoiAdán Pérez-García José Miguel Gasulla Francisco Ortega 2014 A new species of Brodiechelys (Testudines, Pan-Cryptodira) from the Early Cretaceous of Spain: Systematic and palaeobiogeographic implications Acta Palaeontologica Polonica 59 2 333–342 10.4202/app.2012.0059 56200636 free Sp. novValidPérez-García, Gasulla & OrtegaEarly AptianArcillas de Morella FormationSpainA xinjiangchelyid, a species of Brodiechelys.Cheirogaster bacharidisiEvangelos Vlachos Evangelia Tsoukala Joseph Corsini 2014 Cheirogaster bacharidisi, sp. nov., a new species of a giant tortoise from the Pliocene of Thessaloniki (Macedonia, Greece) Journal of Vertebrate Paleontology 34 3 560–575 10.1080/02724634.2013.821409 85589576 https://figshare.com/articles/journal_contribution/1017334 subscription Sp. novValidVlachos, Tsoukala & CorsiniPlioceneGonia FormationGreeceA tortoise, originally described as a species belonging to the genus Cheirogaster. Subsequently, transferred by Pérez-García & Vlachos (2014) to the genus Titanochelon.EodortokaA. Pérez-García J.M. Gasulla F. Ortega 2014 Eodortoka morellana gen. et sp. nov., the first pan-pleurodiran turtle (Dortokidae) defined in the Lower Cretaceous of Europe Cretaceous Research 48 130–138 10.1016/j.cretres.2013.12.004 Gen. et sp. novValidPérez-García, Gasulla & OrtegaEarly Cretaceous (Aptian)Arcillas de Morella FormationSpainA dortokid, a member of the clade Pan-Pleurodira (containing living pleurodirans and all turtles that are more closely related to them than to cryptodirans). The type species is Eodortoka morellana.GobiapaloneGen. et comb. et sp. novValidDanilov et al.Late Cretaceous (Cenomanian to Maastrichtian)Barun Goyot Formation Bayan Shireh Formation Nemegt FormationMongoliaA trionychine trionychid. The type species is "Amyda" orlovi Khosatzky (1976); genus also contains new species Gobiapalone breviplastra. The genus Gobiapalone was considered to be a junior synonym of the genus Kuhnemys by Georgalis & Joyce (2017), though the authors maintained G. orlovi and G. breviplastra as distinct species within the latter genus.Georgios L. Georgalis Walter G. Joyce 2017 A Review of the Fossil Record of Old World Turtles of the Clade Pan-Trionychidae Bulletin of the Peabody Museum of Natural History 58 1 115–208 10.3374/014.058.0106 89732695 http://doc.rero.ch/record/288660/files/joy_rfr.pdf Hylaeochelys kappaAdán Pérez-García Francisco Ortega 2014 A new species of the turtle Hylaeochelys (Eucryptodira) outside its known geographic and stratigraphic ranges of distribution Comptes Rendus Palevol 13 3 183–188 10.1016/j.crpv.2013.10.009 Sp. novValidPérez-García & OrtegaLate Jurassic (Tithonian)Freixial FormationPortugalA basal member of Eucryptodira, a species of Hylaeochelys.Judithemys kranziRobert E. Weems 2014 Paleogene chelonians from Maryland and Virginia PaleoBios 31 1 1–32 https://escholarship.org/uc/item/7253p3tf Sp. nov ValidWeemsPaleocene (early Thanetian)Aquia FormationUnited StatesA member of (likely paraphyletic) group "Macrobaenidae", a species of Judithemys.NemegtemysIgor G. Danilov Ren Hirayama Vladimir B. Sukhanov Shigeru Suzuki Mahito Watabe Natasha S. Vitek 2014 Cretaceous soft-shelled turtles (Trionychidae) of Mongolia: new diversity, records and a revision Journal of Systematic Palaeontology 12 7 799–832 10.1080/14772019.2013.847870 86304259 Gen. et sp. novValidDanilov et al.Late Cretaceous (Maastrichtian)Nemegt FormationMongoliaA cyclanorbine trionychid. The type species is Nemegtemys conflata.OsonachelusFrance de Lapparent de Broin Xabier Murelaga Francesc Farrés Jacint Altimiras 2014 An exceptional cheloniid turtle, Osonachelus decorata nov. gen., nov. sp., from the Eocene (Bartonian) of Catalonia (Spain) Geobios 47 3 111–132 10.1016/j.geobios.2014.02.002 Gen. et sp. novValidde Lapparent de Broin et al.Eocene (late Bartonian)Vic-Manlleu Marls FormationSpainA cheloniid sea turtle. The type species is Osonachelus decorata.Procolpochelys charlestonensisSp. nov ValidWeems & SandersOligoceneUnited StatesA pancheloniid sea turtle; a species of Procolpochelys.200px|RiodevemysAdán Pérez-García Rafael Royo-Torres Alberto Cobos 2015 A new European Late Jurassic pleurosternid (Testudines, Paracryptodira) and a new hypothesis of paracryptodiran phylogeny Journal of Systematic Palaeontology 13 4 351–369 10.1080/14772019.2014.911212 83768369 Gen. et sp. novValidPérez-García, Royo-Torres & CobosLate JurassicSpainA pleurosternid paracryptodiran. The type species is Riodevemys inumbragigas.TacuarembemysDaniel Perea Matías Soto Juliana Sterli Valeria Mesa Pablo Toriño Guillermo Roland Jorge Da Silva 2014 Tacuarembemys kusterae, gen. et sp. nov., a new Late Jurassic–?earliest Cretaceous continental turtle from western Gondwana Journal of Vertebrate Paleontology 34 6 1329–1341 10.1080/02724634.2014.859620 11336/21191 129848452 https://figshare.com/articles/dataset/_i_Tacuarembemys_kusterae_i_gen_et_sp_nov_a_new_Late_Jurassic_earliest_Cretaceous_continental_turtle_from_western_Gondwana/1228037 free Gen. et sp. nov ValidPerea et al.Late Jurassic or earliest CretaceousTacuarembó FormationUruguayA turtle of uncertain phylogenetic placement. The type species is Tacuarembemys kusterae.TitanochelonAdán Pérez-García Evangelos Vlachos 2014 New generic proposal for the European Neogene large testudinids (Cryptodira) and the first phylogenetic hypothesis for the medium and large representatives of the European Cenozoic record Zoological Journal of the Linnean Society 172 3 653–719 10.1111/zoj.12183 86295621 Gen. et comb. novValidPérez-García & VlachosEarly Miocene to early PleistoceneAustria Bulgaria France Germany Greece Portugal Spain Switzerland Turkey Malta?A tortoise; a new genus for "Testudo" bolivari Hernández-Pacheco (1917). Genus also contains "Testudo" eurysternum Gervais (1848–1852), "Testudo" ginsburgi de Broin (1977), "Testudo" vitodurana Biedermann (1862), "Cheirogaster" steinbacheri Karl (1996), "Testudo" leberonensis Depéret (1890), "Testudo" schafferi Szalai (1931), "Testudo" perpiniana Depéret (1885) and "Cheirogaster" bacharidisi Vlachos et al. (2014). Genus might also contain "Testudo" gymnesicus Bate (1914).200px|'Trionyx' baynshirensisSp. novValidDanilov et al.Late Cretaceous (Cenomanian to Santonian)Bayan Shireh FormationMongoliaA trionychine trionychid, a species of Trionyx sensu lato.'Trionyx' dissolutusNatasha S. Vitek Igor G. Danilov 2014 Soft-shelled turtles (Trionychidae) from the Cenomanian of Uzbekistan Cretaceous Research 49 1–12 10.1016/j.cretres.2014.01.004 2152/41076 free Sp. novValidVitek & DanilovLate Cretaceous (Cenomanian)UzbekistanA trionychid, a species of Trionyx sensu lato.'Trionyx' gilbentuensisSp. novValidDanilov et al.Late Cretaceous (Maastrichtian)Nemegt FormationMongoliaA trionychine trionychid, a species of Trionyx sensu lato.'Trionyx' gobiensisSp. novValidDanilov et al.Late Cretaceous (Maastrichtian)Nemegt FormationMongoliaA trionychine trionychid, a species of Trionyx sensu lato.'Trionyx' shiluutulensisSp. novValidDanilov et al.Late Cretaceous (Campanian)MongoliaA trionychine trionychid, a species of Trionyx sensu lato. Archosauromorphs Archosaurs 2014 in archosaur paleontology Basal archosauromorphs Research A redescription of Tasmaniosaurus triassicus is published by Ezcurra (2014).Martín D. Ezcurra 2014 The Osteology of the Basal Archosauromorph Tasmaniosaurus triassicus from the Lower Triassic of Tasmania, Australia PLOS ONE 9 1 e86864 10.1371/journal.pone.0086864 24497988 3907582 2014PLoSO...986864E free A revision of anatomy and phylogenetic relationships of the archosauriform Dorosuchus neoetus is published by Sookias et al. (2014).Roland B. Sookias Andrey G. Sennikov David J. Gower Richard J. Butler 2014 The monophyly of Euparkeriidae (Reptilia: Archosauriformes) and the origins of Archosauria: a revision of Dorosuchus neoetus from the Mid-Triassic of Russia Palaeontology 57 6 1177–1202 10.1111/pala.12110 84365059 free A revision of putative euparkeriids from the Triassic of China is published by Sookias et al. (2014).Roland B. Sookias Corwin Sullivan Jun Liu Richard J. Butler 2014 Systematics of putative euparkeriids (Diapsida: Archosauriformes) from the Triassic of China PeerJ 2 e658 10.7717/peerj.658 25469319 4250070 free New taxa Name Novelty Status Authors Age Unit Location Notes ImagesAenigmastropheusMartín D. Ezcurra Torsten M. Scheyer Richard J. Butler 2014 The Origin and Early Evolution of Sauria: Reassessing the Permian Saurian Fossil Record and the Timing of the Crocodile-Lizard Divergence PLOS ONE 9 2 e89165 10.1371/journal.pone.0089165 24586565 3937355 2014PLoSO...989165E free Gen. et sp. nov ValidEzcurra, Scheyer & BulterLate Permian (Wuchiapingian)Usili FormationTanzaniaA non-archosauriform archosauromorph, probably a protorosaurian. The type species is Aenigmastropheus parringtoni.200px|Garjainia madibaDavid J. Gower P. John Hancox Jennifer Botha-Brink Andrey G. Sennikov Richard J. Butler 2014 A New Species of Garjainia Ochev, 1958 (Diapsida: Archosauriformes: Erythrosuchidae) from the Early Triassic of South Africa PLOS ONE 9 11 e111154 10.1371/journal.pone.0111154 25386937 4227673 2014PLoSO...9k1154G free Sp. nov ValidGower et al.Early Triassic (late Olenekian)Burgersdorp FormationSouth AfricaAn erythrosuchid archosauriform, a species of Garjainia.200px|Hyperodapedon tikiensisDebarati Mukherjee Sanghamitra Ray 2014 A new Hyperodapedon (Archosauromorpha, Rhynchosauria) from the Upper Triassic of India: implications for rhynchosaur phylogeny Palaeontology 57 6 1241–1276 10.1111/pala.12113 86195905 free Sp. nov ValidMukherjee & RayLate TriassicTiki FormationIndiaA rhynchosaur, a species of Hyperodapedon.Proterosuchus goweriMartín D. Ezcurra Richard J. Butler 2015 Taxonomy of the proterosuchid archosauriforms (Diapsida: Archosauromorpha) from the earliest Triassic of South Africa, and implications for the early archosauriform radiation Palaeontology 58 1 141–170 10.1111/pala.12130 84348015 Sp. nov ValidEzcurra & ButlerEarly TriassicSouth AfricaA proterosuchid archosauriform, a species of Proterosuchus.PseudochampsaMaría Jimena Trotteyn Martín D. Ezcurra 2014 Osteology of Pseudochampsa ischigualastensis gen. et comb. nov. (Archosauriformes: Proterochampsidae) from the Early Late Triassic Ischigualasto Formation of Northwestern Argentina PLOS ONE 9 11 e111388 10.1371/journal.pone.0111388 25426846 4245112 2014PLoSO...9k1388T free Gen. et comb. novValidTrotteyn & EzcurraLate Triassic (late Carnian or earliest Norian)Ischigualasto FormationArgentinaA proterochampsid archosauriform; a new genus for "Chanaresuchus" ischigualastensis Trotteyn, Martínez & Alcober (2012).200px| Other reptiles Name Novelty Status Authors Age Unit Location Notes ImagesAbyssomedonMark J. MacDougall Robert R. Reisz 2014 The first record of a nyctiphruretid parareptile from the Early Permian of North America, with a discussion of parareptilian temporal fenestration Zoological Journal of the Linnean Society 172 3 616–630 10.1111/zoj.12180 Gen. et sp. novValidMacdougall & ReiszEarly PermianGarber FormationUnited StatesA nyctiphruretid parareptile. The type species is Abyssomedon williamsi.200px|AtopodentatusLong Cheng Xiao-Hong Chen Qing-Hua Shang Xiao-Chun Wu amp 2014 A new marine reptile from the Triassic of China, with a highly specialized feeding adaptation Naturwissenschaften 101 3 251–259 10.1007/s00114-014-1148-4 24452285 2014NW....101..251C 7770583 Gen. et sp. novValidCheng et al.Middle Triassic (Anisian)Guanling FormationChinaA marine reptile, probably a relative of sauropterygians. The type species is Atopodentatus unicus.200px| Cartorhynchus Ryosuke Motani Da-Yong Jiang Guan-Bao Chen Andrea Tintori Olivier Rieppel Cheng Ji Jian-Dong Huang amp 2014 A basal ichthyosauriform with a short snout from the Lower Triassic of China Nature 517 485–488 10.1038/nature13866 Gen. et sp. novValidMotani et al.Lower TriassicNanlinghu FormationChinaA basal member of Ichthyosauriformes. The type species is C. lenticarpus. Published online in 2014; the final version of the article naming it was published in 2015.200px|Delorhynchus cifelliiRobert R. Reisz Mark J. Macdougall Sean P. Modesto 2014 A new species of the parareptile genus Delorhynchus, based on articulated skeletal remains from Richards Spur, Lower Permian of Oklahoma Journal of Vertebrate Paleontology 34 5 1033–1043 10.1080/02724634.2013.829844 128459194 Sp. novValidReisz, Macdougall & ModestoEarly PermianGarber FormationUnited StatesA parareptile relative of lanthanosuchoids, a species of Delorhynchus.200px|EohupehsuchusXiao-hong Chen Ryosuke Motani Long Cheng Da-yong Jiang Olivier Rieppel amp 2014 A Small Short-Necked Hupehsuchian from the Lower Triassic of Hubei Province, China PLOS ONE 9 12 e115244 10.1371/journal.pone.0115244 25517113 4269458 2014PLoSO...9k5244C free Gen. et sp. novValidChen et al.Early TriassicJialingjiang FormationChinaA hupehsuchian. The type species is Eohupehsuchus brevicollis.HoraffiaNicole Klein Hans Hagdorn amp 2014 Humerus morphology and histology of a new marine reptile (Diapsida) from the Muschelkalk-Keuper-Grenzbonebed (Middle Triassic, Ladinian) of Southwest Germany Palaeodiversity 7 23–38 http://www.palaeodiversity.org/pdf/07/02Palaeodiversity_7-14_Klein_3.pdf Gen. et sp. novValidKlein & HagdornMiddle Triassic (Ladinian)Erfurt FormationGermanyA marine diapsid reptile of uncertain phylogenetic placement. The type species is Horaffia kugleri.Largocephalosaurus qianensisChun Li Da-Yong Jiang Long Cheng Xiao-Chun Wu Olivier Rieppel amp 2014 A new species of Largocephalosaurus (Diapsida: Saurosphargidae), with implications for the morphological diversity and phylogeny of the group Geological Magazine 151 1 100–120 10.1017/S001675681300023X 2014GeoM..151..100L 84534618 Sp. novValidLi et al.TriassicGuanling FormationChinaA member of Saurosphargidae, a species of Largocephalosaurus.ParahupehsuchusXiao-hong Chen Ryosuke Motani Long Cheng Da-yong Jiang Olivier Rieppel amp 2014 A Carapace-Like Bony 'Body Tube' in an Early Triassic Marine Reptile and the Onset of Marine Tetrapod Predation PLOS ONE 9 4 e94396 10.1371/journal.pone.0094396 24718682 3981804 2014PLoSO...994396C free Gen. et sp. novValidChen et al.Early TriassicJialingjiang FormationChinaA hupehsuchian. The type species is Parahupehsuchus longus. References Category:2014 in paleontology