2012 in paleoentomology

none 2012 in arthropod paleontology 2012 in paleontology 2012 paleoentomology paleontology paleobotany arthropod paleontology paleoichthyology paleomalacology paleomammalogy reptile paleontology archosaur paleontology This list of 2012 in paleoentomology records new fossil insect taxa that were described during the year, as well as documents significant paleoentomology discoveries and events which occurred during that year. 3 Blattodea Name Novelty Status Authors Age Unit Location Notes ImagesArchimesoblatta shivaMichael S. Engel Ricardo Pérez-de La Fuente 2012 A New Species of Roach from the Jurassic of India (Blattaria: Mesoblattinidae) Journal of the Kansas Entomological Society 85 1 1–4 10.2317/JKES110524.1 86567655 Sp. novValidEngel & Pérez-de La FuenteMiddle JurassicKota FormationIndiaA mesoblattinid cockroachArcofuziaDandan Wei Chungkun Shih Dong Ren 2012 Arcofuzia cana gen. et sp. n. (Insecta, Blattaria, Fuziidae) from the Middle Jurassic sediments of Inner Mongolia, China Zootaxa 3597 25–32 https://www.mapress.com/zootaxa/2012/f/z03597p032f.pdf 10.11646/zootaxa.3597.1.3 Gen. et sp. novValidWei, Shih & RenMiddle JurassicJiulongshan FormationChinaA fuziid cockroach. The type species is Arcofuzia cana.Cariblattoides labandeiraiVršanský P. Vidlička L. Čiampor Jr Marsh F. 2012 Derived, still living cockroach genus Cariblattoides (Blattida: Blattellidae) from the Eocene sediments of Green River in Colorado, USA Insect Science 19 2 143–152 10.1111/j.1744-7917.2010.01390.x 2012InsSc..19..143V 32630827 Sp. novValidVršanský et al.EoceneBridgerianGreen River FormationLake UnitaUnited StatesColoradoA Pseudophyllodromiine wood cockroachDivocinaJun-Hui Liang Peter Vršanský Dong Ren 2012 Variability and symmetry of a Jurassic nocturnal predatory cockroach (Blattida: Raphidiomimidae) Revista Mexicana de Ciencias Geológicas 29 2 411–421 http://satori.geociencias.unam.mx/29-2/%2809%29Liang.pdf 2013-01-11 2014-02-23 https://web.archive.org/web/20140223051546/http://satori.geociencias.unam.mx/29-2/%2809%29Liang.pdf dead Gen. et sp. nov.ValidLiang, Vršanský & RenMiddle Jurassic (Bathonian)Jiulongshan FormationChinaA raphidiomimid cockroach. The type species is Divocina noci.EntropiaPeter Vršanský Jun-Hui Liang Dong Ren 2012 Malformed cockroach (Blattida: Liberiblattinidae) in the Middle Jurassic sediments from China Oriental Insects 46 1 12–18 10.1080/00305316.2012.675482 85209650 Gen. et sp. novValidVršanský, Liang & RenMiddle Jurassic (Bathonian)ChinaA liberiblattinid cockroach. The type species is Entropia initialis.GraciliblattaJun-Hui Liang Wei-Long Huang Dong Ren 2012 Graciliblatta bella gen. et sp. n. — a rare carnivorous cockroach (Insecta, Blattida, Raphidiomimidae) from the Middle Jurassic sediments of Daohugou in Inner Mongolia, China Zootaxa 3449 62–68 https://www.mapress.com/zootaxa/2012/f/z03449p068f.pdf 10.11646/zootaxa.3449.1.4 Gen. et sp. novValidLiang, Huang & RenMiddle JurassicJiulongshan FormationChinaA raphidiomimid cockroach. The type species is Graciliblatta bella.MutoviaPeter Vršanský Danil Aristov 2012 Enigmatic Late Permian cockroaches from Isady, Russia (Blattida: Mutoviidae fam. n.) Zootaxa 3247 19–31 https://www.mapress.com/zootaxa/2012/f/z03247p031f.pdf 10.11646/zootaxa.3247.1.2 Gen. et sp. novValidVršanský & AristovLate Permian RussiaA mutoviid cockroach. The type species is Mutovia intercalaria.Parvifuzia peregrinaDandan Wei Junhui Liang Dong Ren 2012 A new species of Fuziidae (Insecta, Blattida) from the Inner Mongolia, China ZooKeys 217 53–61 https://www.pensoft.net/journals/zookeys/article/3508/a-new-species-of-fuziidae-insecta-blattida-from-the-inner-mongolia-china 10.3897/zookeys.217.3508 22977342 3433702 free 2012ZooK..217...53D Sp. novValidWei, Liang & RenMiddle JurassicJiulongshan FormationChinaA fuziid cockroach, a species of Parvifuzia.QilianiblattaZhijun Zhang Joerg W. Schneider Youchong Hong 2013 The most ancient roach (Blattodea): a new genus and species from the earliest Late Carboniferous (Namurian) of China, with a discussion of the phylomorphogeny of early blattids Journal of Systematic Palaeontology 11 1 27–40 10.1080/14772019.2011.634443 2013JSPal..11...27Z 85244513 Sp. novValidZhang, Schneider & HongCarboniferous (earliest Pennsylvanian)ChinaA cockroach. The type species is Qilianiblatta namurensis.StegoblattaL.N. Anisyutkin C. Gröhn 2012 New cockroaches (Dictyoptera: Blattina) from Baltic Amber, with description of a new genus and species: Stegoblatta irmgardgroehni Proceedings of the Zoological Institute of the Russian Academy of Sciences 316 3 193–202 https://www.zin.ru/journals/trudyzin/doc/vol_316_3/TZ_316_3_Anisyutkin.pdfGen. et sp. novValidAnisyutkin & GröhnEoceneBaltic amberEuropeA blaberid cockroach. The type species is Stegoblatta irmgardgroehni. Coleoptera Name Novelty Status Authors Age Unit Location Notes ImagesAbrocar concavusSteve R. Davis Michael S. Engel Andrei Legalov Dong Ren 2013 Weevils of the Yixian Formation, China (Coleoptera: Curculionoidea): phylogenetic considerations and comparison with other Mesozoic faunas Journal of Systematic Palaeontology 11 4 399–429 10.1080/14772019.2012.691906 2013JSPal..11..399D 84192417 Sp. novValidDavis et al.Early CretaceousYixian FormationChinaA weevil, a species of Abrocar.Abrocar relicinusSp. novValidDavis et al.Early CretaceousYixian FormationChinaA weevil, a species of Abrocar.AbrohadeocoleodesJingjing Tan Dong Ren Chungkun Shih Xingke Yang 2013 New schizophorid fossils from China and possible evolutionary scenarios for Jurassic archostematan beetles Journal of Systematic Palaeontology 11 1 47–62 10.1080/14772019.2011.637515 2013JSPal..11...47T 83526884 Gen. et 4 sp. novValidTan et al.Middle JurassicJiulongshan FormationChinaA schizophorid archostematan beetle. Genus contains four species: A. eurycladus, A. ooideus, A. nii and A. patefactus.AlloioscarabaeusMing Bai Dirk Ahrens Xing-Ke Yang Dong Ren 2012 New fossil evidence of the early diversification of scarabs: Alloioscarabaeus cheni (Coleoptera: Scarabaeoidea) from the Middle Jurassic of Inner Mongolia, China Insect Science 19 2 159–171 10.1111/j.1744-7917.2011.01460.x free 2012InsSc..19..159B Gen. et sp. novValidBai, Ren & Yang in Bai et al.Middle JurassicJiulongshan FormationChinaAn alloioscarabaeid scarabaeoid beetle. The type species is Alloioscarabaeus cheni.AlphitopsisA. G. Kirejtshuk M. V. Nabozhenko A. Nel 2012 First Mesozoic representative of the subfamily Tenebrioninae (Coleoptera, Tenebrionidae) from the Lower Cretaceous of Yixian (China, Liaoning) Entomological Review 92 1 97–100 10.1134/S0013873812010101 2012EntRv..92...97K 14365498 Gen. et sp. novValidKirejtshuk, Nabozhenko & NelEarly CretaceousYixian FormationChinaA tenebrionine darkling beetle. The type species is Alphitopsis initialis.Anotylus archaicusYanli Yue György Makranczy Dong Ren 2012 A Mesozoic Species of Anotylus (Coleoptera, Staphylinidae, Oxytelinae) from Liaoning, China, With the Earliest Evidence of Sexual Dimorphism In Rove Beetles Journal of Paleontology 86 3 508–512 http://jpaleontol.geoscienceworld.org/content/86/3/508.abstract 10.1666/11-032.1 2012JPal...86..508Y 67830565 subscription sp novValidYue, Makranczy & RenEarly Cretaceous Yixian FormationChinaAn oxyteline rove beetle, a species of Anotylus.AphebodactylaStylianos Chatzimanolis Molly E. Cashion Michael S. Engel Zachary H. Falin 2012 A new genus of Ptilodactylidae (Coleoptera: Byrrhoidea) in mid-Cretaceous amber from Myanmar (Burma) Geodiversitas 34 3 569–574 10.5252/g2012n3a6 2012Geodv..34..569C 85168129 http://sciencepress.mnhn.fr/en/periodiques/geodiversitas/34/3/nouveau-genre-de-ptilodactylidae-coleoptera-byrrhoidea-de-l-ambre-albien-terminal-du-myanmar-birmanie Gen. et sp. novValidChatzimanolis et al.CretaceousBurmese amberMyanmarA ptilodactylid byrrhoid beetle. The type species is Aphebodactyla rhetine.Aploceble (Chalcoaploceble) viridiaeneusS.E. Tshernyshev 2012 Aploceble (Chalcoaploceble) viridiaeneus Tshernyshev — new subgenus and species of dasytid beetles (Coleoptera, Dasytidae) in Baltic Amber Euroasian Entomological Journal 11 3 213–281 https://www.researchgate.net/publication/288653983 Subgen. et sp. novValidTshernyshevEoceneBaltic amberRussia (Kaliningrad Oblast)A melyrid beetle belonging to the subfamily Dasytinae, a subgenus and species of Aploceble.ApticaxAndrea Schomann Alexey Solodovnikov 2012 A new genus of Staphylinidae (Coleoptera) from the Lower Cretaceous: the first fossil rove beetles from the Southern Hemisphere Systematic Entomology 37 2 379–386 10.1111/j.1365-3113.2011.00615.x 2012SysEn..37..379S 83673133 Gen. et 2 sp. novValidSchomann & SolodovnikovEarly Cretaceous (Aptian–Albian)Crato FormationBrazilA rove beetle of uncertain phylogenetic placement. The type species is A. volans; genus also includes A. solidus.ArchaeosciaphilusA. A. Legalov 2012 New curculionoid beetles (Coleoptera: Curculionoidea) from the Baltic amber Paleontological Journal 46 3 262–272 10.1134/S0031030112030094 2012PalJ...46..262L 85795686 Gen. et sp. novValidLegalovEoceneEuropeA curculionid beetle found in Baltic amber. The type species is Archaeosciaphilus marshalli.Attagenus yantarnyiJiří Háva Andris Bukejs 2012 Attagenus yantarnyi sp. nov., a new species from Baltic amber (Coleoptera: Dermestidae) Baltic Journal of Coleopterology 12 2 155–158 Sp. novValidHáva & BukejsLate EocenePrussian FormationRussia (Kaliningrad Oblast)A dermestid beetle found in Baltic amber, a species of Attagenus.Baltocar groehniAlexander Riedel Tomy dos Santos Rolo Angelica Cecilia Thomas van de Kamp 2012 Sayrevilleinae Legalov, a newly recognised subfamily of fossil weevils (Coleoptera, Curculionoidea, Attelabidae) and the use of synchrotron microtomography to examine inclusions in amber Zoological Journal of the Linnean Society 165 4 773–794 10.1111/j.1096-3642.2012.00825.x free Sp. novValidRiedel in Riedel et al.EoceneBaltic amberRussia(Kaliningrad Oblast)A weevil belonging to the family Attelabidae.Baltocar hoffeinsorumSp. novValidRiedel in Riedel et al.EoceneBaltic amberRussia(Kaliningrad Oblast)A weevil belonging to the family Attelabidae.Baltocar subnudusSp. novValidRiedel in Riedel et al.EoceneBaltic amberRussia(Kaliningrad Oblast)A weevil belonging to the family Attelabidae.Cartodere (Aridius) succinobalticaAndris Bukejs Wolfgang H. Rücker Alexander G. Kirejtshuk 2012 Cartodere (Aridius) succinobaltica sp. nov. (Coleoptera: Latridiidae) from Baltic amber Latridiidae 9 25–28 https://www.zin.ru/animalia/coleoptera/pdf/bukejs_et_al_9_2012_bernstein.pdf Sp. novValidBukejs, Rücker & KirejtshukLate EoceneEuropeA latridiid beetle found in Baltic amber, a species of Cartodere.Catops perkovskyiMichel Perreau 2012 Description of a new genus and two new species of Leiodidae (Coleoptera) from Baltic amber using phase contrast synchrotron X-ray microtomography Zootaxa 3455 81–88 https://www.mapress.com/zootaxa/2012/f/zt03455p088.pdf 10.11646/zootaxa.3455.1.5 free Sp. novValidPerreauEoceneRussia (Kaliningrad Oblast)A leiodid beetle found in Baltic amber, a species of Catops.Conapium (Palaeoconapion) alleniAndrei A. Legalov 2012 New and little known Apioninae (Coleoptera, Brentidae) in Eocene Baltic amber Euroasian Entomological Journal 11 3 219–222 https://elibrary.ru/item.asp?id=17892054 Subgen. et sp. novValidLegalovEoceneBaltic amberRussia (Kaliningrad Oblast)A beetle belonging to the family Brentidae, a subgenus and species of Conapium.CrassisorusG.V. Nikolajev B. Wang H. Zhang 2012 A new genus of the family Hybosoridae (Coleoptera, Scarabaeoidea) from the Yixian Lower Cretaceous Formation in China Euroasian Entomological Journal 11 6 503–505 https://elibrary.ru/item.asp?id=18428007 Gen. et sp. novValidNikolajev, Wang & ZhangEarly CretaceousYixian FormationChinaA member of the family Hybosoridae. The type species is C. fractus.CretohypnaZhuo Yan Georgiy V. Nikolajev Dong Ren 2012 A new, well-preserved genus and species of fossil Glaphyridae (Coleoptera, Scarabaeoidea) from the Mesozoic Yixian Formation of Inner Mongolia, China ZooKeys 241 67–75 10.3897/zookeys.241.3262 23372414 3559105 free 2012ZooK..241...67Y Gen. et sp. novValidYan, Nikolajev & RenEarly CretaceousYixian FormationChinaA glaphyrid beetle. The type species is Cretohypna cristata.CretotanaosAndrei A. Legalov 2012 The oldest Brentidae and Curculionidae (Coleoptera: Curculionoidea) from the Aptian of Bon-Tsagaan Historical Biology: An International Journal of Paleobiology 26 1 6–15 10.1080/08912963.2012.751103 85027770 Gen. et sp. novValidLegalovEarly Cretaceous (Aptian)MongoliaAn apionine brentid weevil. Type species is Cretotanaos bontsaganensis.Cryptophagus harenusG. Yu. Lyubarsky E. E. Perkovsky 2012 The First Eocene Species of the Genus Cryptophagus (Coleoptera, Clavicornia, Cryptophagidae) Vestnik Zoologii 46 1 83–87 https://www.v-zool.kiev.ua/tocs/e46-1.htm 10.2478/v10058-012-0007-z free Sp. novValidLyubarsky & PerkovskyLate EoceneUkraineA cryptophagid beetle found in Rovno amber, a species of Cryptophagus.ElectotretaGen. et sp.ValidKazantsevEocene LithuaniaA firefly. The type species is Electotreta rasnitsyni.ElectromethesSergey V. Kazantsev 2012 New omethid and lampyrid taxa from the Baltic Amber (Insecta: Coleoptera) Zootaxa 3186 59–63 https://www.mapress.com/zootaxa/2012/f/z03186p063f.pdf 10.11646/zootaxa.3186.1.5 Gen. et sp. novValidKazantsevEocene LithuaniaAn omethid beetle. The type species is Electromethes alleni.ElectropteronSergey V. Kazantsev 2012 A New Lycid Genus from the Dominican Amber (Insecta, Coleoptera, Lycidae, Leptolycinae, Leptolycini) Psyche: A Journal of Entomology 2012 1–3 10.1155/2012/982141 free Gen. et sp. novValidKazantsevOligocene Dominican RepublicA lycid. The type species is E. avus.Elodes beigeliB. Klausnitzer 2012 Neue Arten der Familie Scirtidae (Coleoptera) aus Baltischem Bernstein (Teil 3) (172. Beitrag zur Kenntnis der Scirtidae) Linzer Biologische Beiträge 44 1 313–318 https://www.landesmuseum.at/pdf_frei_remote/LBB_0044_1_0313-0318.pdf Sp. novValidKlausnitzerEoceneEuropeA scirtid beetle found in Baltic amber, a species of Elodes.Elodes mysticopalpalisSp. novValidKlausnitzerEoceneEuropeA scirtid beetle found in Baltic amber, a species of Elodes.Enicmus adrianaeAndris Bukejs Hans-Peter Reike Wolfgang H. Rücker 2012 Enicmus adrianae sp. nov. - a new scavenger beetle (Coleoptera: Latridiidae) from Baltic amber Baltic Journal of Coleopterology 12 2 149–154 http://www.bjc.sggw.pl/viewart.php?bookid=23&artid=449 Sp. novValidBukejs, Reike & RückerLate EoceneRussia (Kaliningrad Oblast)A member of Latridiidae found in Baltic amber, a species of Enicmus.EocenorhynchitesGen. et sp. novValidLegalovEoceneEuropeA rhynchitid beetle found in Baltic amber. The type species is Eocenorhynchites vossi.Eodromeus daohugouensisBo Wang Haichun Zhang Alexandr G. Ponomarenko 2012 Mesozoic Trachypachidae (Insecta: Coleoptera) from China Palaeontology 55 2 341–353 10.1111/j.1475-4983.2012.01128.x 2012Palgy..55..341W free sp novValidWang, Zhang and PonomarenkoMiddle JurassicDaohugou BedsChinaA trachypachid, a species of Eodromeus.Eodromeus robustussp novValidWang, Zhang and PonomarenkoMiddle JurassicDaohugou BedsChinaA trachypachid, a species of Eodromeus.EoluciolaSergey V. Kazantsev 2012 A new Luciolinae firefly (Coleoptera: Lampyridae) from the Baltic Amber Russian Entomological Journal 21 3 319–320 https://kmkjournals.com/upload/PDF/REJ/21/ent21_3%20319_320%20Kazantsev%20for%20Inet.pdf 10.15298/rusentj.21.3.08 free Gen. et sp. novValidKazantsevEoceneEurope (Baltic Sea coast)A lucioline firefly found in Baltic amber. The type species is Eoluciola varang.EuroleptochromusPaweł Jałoszyński 2012 Description of Euroleptochromus gen.n. (Coleoptera, Staphylinidae, Scydmaeninae) from Baltic amber, with discussion of biogeography and mouthpart evolution within Clidicini Systematic Entomology 37 2 346–359 10.1111/j.1365-3113.2012.00623.x 2012SysEn..37..346J 222203806 Gen. et sp. novValidJałoszyńskiEocene LithuaniaA rove beetle found in Lithuanian Eocene Baltic amber. The type species is Euroleptochromus sabathi.Helophorus (Mesosperchus) inceptivusMartin Fikáček Alexander Prokin Robert B. Angus Alexander Ponomarenko Yanli Yue Dong Ren Jakub Prokop 2012 Revision of Mesozoic fossils of the helophorid lineage of the superfamily Hydrophiloidea (Coleoptera: Polyphaga) Acta Entomologica Musei Nationalis Pragae 52 1 89–127 https://www.aemnp.eu/data/article-1355/1336-52_1_89.pdf Sp. novValidFikáček et al.Late JurassicMongoliaA hydrophiloid beetle, a species of Helophorus.Helophorus (Mesosperchus) yixianusSp. novValidFikáček et al.Jurassic-Cretaceous boundaryYixian FormationChinaA hydrophiloid beetle, a species of Helophorus.Hydroscapha? jeholensisChenyang Cai Andrew E. Z. Short Diying Huang 2012 The first skiff beetle (Coleoptera: Myxophaga: Hydroscaphidae) from Early Cretaceous Jehol Biota Journal of Paleontology 86 1 116–119 10.1666/11-050.1 2012JPal...86..116C 140170420 Sp. novValidCai, Short & HuangEarly CretaceousYixian FormationChinaA member of the family Hydroscaphidae.JuroglypholomaChenyang Cai Diying Huang Margaret K. Thayer Alfred F. Newton 2012 Glypholomatine Rove Beetles (Coleoptera: Staphylinidae): a Southern Hemisphere Recent Group Recorded from the Middle Jurassic of China Journal of the Kansas Entomological Society 85 3 239–244 10.2317/JKES120531.1 86755283 Gen. et sp. novValidCai et al.Middle JurassicDaohugou BedsChinaA glypholomatine rove beetle. The type species is Juroglypholoma antiquum.Kararhynchus gratsheviSpeciesValidLegalovMiddle-Late Jurassic Karabastau FormationKazakhstanAn obrieniid beetle, a species of Kararhynchus.Kararhynchus jurassicusSpeciesValidLegalovMiddle-Late Jurassic Karabastau FormationKazakhstanAn obrieniid beetle, a species of Kararhynchus.LaetopsiaGen. et comb. novValidFikáček et al.Early CretaceousChina Mongolia RussiaA hydrophiloid beetle; a new genus for "Hydrophilopsia" baissensis Ponomarenko (1987), "H." bontsaganica Prokin (2009), "H." hydraenoides Prokin, Ren & Fikáček (2010), "H." mongolica Ponomarenko (1987) and "H." shatrovskiyi Prokin, Ren & Fikáček (2010).Lasiosyne cataphractaE. V. Yan 2012 New beetle species of the genus Lasiosyne (Coleoptera, Lasiosynidae) from the Late Jurassic and Early Cretaceous of Russia and Mongolia Paleontological Journal 46 6 589–600 10.1134/S0031030112060111 2012PalJ...46..589Y 84507083 Sp. novValidYanEarly CretaceousMongoliaA lasiosynid beetle, a species of Lasiosyne.Lasiosyne decoraSp. novValidYanEarly CretaceousRussiaA lasiosynid beetle, a species of Lasiosyne.Lasiosyne insculptaSp. novValidYanLate JurassicMongoliaA lasiosynid beetle, a species of Lasiosyne.Lasiosyne longitarsaSp. novValidYanLate JurassicMongoliaA lasiosynid beetle, a species of Lasiosyne.Lasiosyne punctataSp. novValidYanEarly CretaceousMongoliaA lasiosynid beetle, a species of Lasiosyne.Lasiosyne shartegiensisSp. novValidYanLate JurassicMongoliaA lasiosynid beetle, a species of Lasiosyne.LeehermaniaStylianos Chatzimanolis David A. Grimaldi Michael S. Engel Nicholas C. Fraser 2012 Leehermania prorova, the earliest staphyliniform beetle, from the late Triassic of Virginia American Museum Novitates 3761 1–28 10.1206/3761.2 2246/6393 86429652 https://zenodo.org/record/4597978 Gen. et sp. novValidChatzimanolis, Grimaldi & Engel in Chatzimanolis et al.Late Triassic (late Carnian or early Norian)Cow Branch FormationUnited StatesA beetle of uncertain phylogenetic placement, might be the earliest known rove beetle or a member of Myxophaga related to the family Hydroscaphidae.Martin Fikáček Rolf G. Beutel Chenyang Cai John F. Lawrence Alfred F. Newton Alexey Solodovnikov Adam Ślipiński Margaret K. Thayer Shûhei Yamamoto 2019 Reliable placement of beetle fossils via phylogenetic analyses – Triassic Leehermania as a case study (Staphylinidae or Myxophaga?) Systematic Entomology 45 175–187 10.1111/syen.12386 The type species is Leehermania prorova.Lithohypna lepticephalaG. V. Nikolajev D. Ren 2012 New species of the genus Lithohypna Nikolajev, Wang et Zhang, 2011 (Coleoptera, Scarabaeoidea, Glaphyridae) from the Yixian Formation, China Euroasian Entomological Journal 11 3 209–211 https://elibrary.ru/item.asp?id=17892052 Sp. novValidNikolajev & RenEarly CretaceousYixian FormationChinaA beetle belonging to the family Glaphyridae, a species of Lithohypna.Lithohypna longulaSp. novValidNikolajev & RenEarly CretaceousYixian FormationChinaA beetle belonging to the family Glaphyridae, a species of Lithohypna.Lithohypna tuberculataSp. novValidNikolajev & RenEarly CretaceousYixian FormationChinaA beetle belonging to the family Glaphyridae, a species of Lithohypna.Lithohypna yuxianaSp. novValidNikolajev & RenEarly CretaceousYixian FormationChinaA beetle belonging to the family Glaphyridae, a species of Lithohypna.Macratria alleniDmitry Telnov 2012 Macratriinae (Coleoptera: Anthicidae) of the Baltic amber Latvijas Entomologs 51 27–39 http://leb.daba.lv/Macratriinae.pdfSp. novValidTelnovLate EoceneLatvia or LithuaniaA macratriine anthicid beetle found in Baltic amber, a species of Macratria.Melanapion (Melanapionoides) wanatiSubgen. et sp. novValidLegalovEoceneBaltic amberRussia (Kaliningrad Oblast)A beetle belonging to the family Brentidae, a subgenus and species of Melanapion.Menopraesagus oryziformisSp. novValidTan et al.Middle JurassicJiulongshan FormationChinaA schizophorid archostematan beetle, a species of Menopraesagus.Nephus subcircularisA.G. Kirejtshuk A. Nel 2012 The oldest representatives of the family Coccinellidae(Coleoptera: Polyphaga) from the Lowermost Eocene Oise amber (France) Zoosystematica Rossica 21 1 131–144 10.31610/zsr/2012.21.1.131 https://www.zin.ru/journals/zsr/pdf/2012-1/ZR_21%281%29_Kirejtshuk_Nel.pdfSp. novValidKirejtshuk & NelEarliest EoceneFranceA scymnine coccinellid beetle found in Oise amber, as species of NephusNotocupes liniSp. novValidPonomarenko et al.Early JurassicChinaOriginally described as a species of Notocupes; Kirejtshuk (2020) transferred this species to the genus Zygadenia.OlemehliaJan Batelka 2012 Olemehlia krali, a new genus and species of Ripidiinae from Baltic amber (Coleoptera: Ripiphoridae) Annales de la Société Entomologique de France Nouvelle Série 47 3–4 361–364 10.1080/00379271.2011.10697729 83868089 Gen. et sp. novJunior homonymBatelkaEocene (Lutetian)Baltic amberLithuaniaA member of the family Ripiphoridae belonging to the subfamily Ripidiinae. The type species is O. krali. The generic name is preoccupied by Olemehlia Holzschuh (2011); Batelka (2017) coined a replacement name Olemehliella.Jan Batelka 2017 A replacement name for the Baltic amber ripidiine genus Olemehlia (Coleoptera: Ripiphoridae) Novitates Paleoentomologicae 20 20 1–2 10.17161/np.v0i20.6543 free Omma delicataJingjing Tan Yongjie Wang Dong Ren Xingke Yang 2012 New fossil species of ommatids (Coleoptera: Archostemata) from the Middle Mesozoic of China illuminating the phylogeny of Ommatidae BMC Evolutionary Biology 12 1 113 10.1186/1471-2148-12-113 22776212 3518168 free 2012BMCEE..12..113T Sp. novTan et al.Middle JurassicJiulongshan FormationChinaA species of OmmaOrchesia (Orchestera) turkiniVitaly I. Alekseev Andris Bukejs 2012 A new species of Orchesia Latreille, 1807 from Baltic amber (Coleoptera: Melandryidae) Genus 23 4 571–576 http://www.biol.uni.wroc.pl/cassidae/Alekseev%20&%20Bukejs_A%20new%20species%20of%20Orchesia%20from%20Baltic%20Amber.pdf 2018-11-18 https://web.archive.org/web/20160304025826/http://www.biol.uni.wroc.pl/cassidae/Alekseev%20%26%20Bukejs_A%20new%20species%20of%20Orchesia%20from%20Baltic%20Amber.pdf 2016-03-04 dead Sp. novValidAlekseev & BukejsLate EocenePrussian FormationRussia (Kaliningrad Oblast)A melandryid beetle found in Baltic amber, a species of Orchesia.PalaeoerirhinusGen. et 4 sp. novValidLegalovEarly Cretaceous (Aptian)MongoliaA curculionid weevil. Type species is Palaeoerirhinus ponomarenkoi; genus also contains new species P. latus, P. thompsoni and P. longirostris.PalaeometrioxenaGen. et sp. novJunior synonymLegalovEoceneBaltic amberEurope (Baltic Sea region)A belid. The type species is Palaeometrioxena zherikhini. Legalov (2019) considered the genus Palaeometrioxena to be a junior synonym of the genus Archimetrioxena, and transferred the species P zherikhini to the latter genus.PareuryommaNom. et 2 sp. novTan et al.Early CretaceousYixian FormationChinaA member of the family Ommatidae; a replacement name for Euryomma Tan et al. (2006). The type species is "Euryomma" tylodes Tan et al. (2006); genus also includes new species P. cardiobasis. Additional new species described by Tan et al., P. ancistrodonta from the Middle Jurassic Jiulongshan Formation, was subsequently transferred to the genus Omma by Kirejtshuk (2020).Pleocoma dolichophyllaG. V. Nikolajev D. Ren 2012 The earliest known species of the genus Pleocoma LeConte (Coleoptera, Scarabaeoidea, Pleocomidae) from the Mesozoic of China Paleontological Journal 46 5 495–498 10.1134/S0031030112050061 2012PalJ...46..495N 84410917 Sp. novValidNikolajev & RenEarly CretaceousYixian FormationChinaA rain beetle, a species of Pleocoma.ProsinodendronMing Bai Dong Ren Xingke Yang 2012 Prosinodendron krelli from the Yixian Formation, China: A missing link among Lucanidae, Diphyllostomatidae and Passalidae (Coleoptera: Scarabaeoidea) Cretaceous Research 34 334–339 10.1016/j.cretres.2011.11.017 2012CrRes..34..334B Gen. et sp. novValidBai, Ren, & YangBarremian to early Aptian Yixian FormationChinaA stag beetle. The type species is Prosinodendron krelli.ProsolieriusMargaret K. Thayer Alfred F. Newton Stylianos Chatzimanolis 2012 Prosolierius, a new mid-Cretaceous genus of Solieriinae (Coleoptera: Staphylinidae) with three new species from Burmese amber Cretaceous Research 34 124–134 10.1016/j.cretres.2011.10.010 2012CrRes..34..124T Gen. et 3 sp. novValidThayer, Newton, & ChatzimanolisBarremian to AlbianLebanon MyanmarA rove beetle.PseudobrieniaA. A. Legalov 2012 New Obrieniidae from the Jurassic of Kazakhstan (Coleoptera, Obrienioidea) Paleontological Journal 46 1 73–78 10.1134/S0031030112010091 2012PalJ...46...73L 86433414 Gen. et sp. novValidLegalovMiddle-Late Jurassic Karabastau FormationKazakhstanAn obrieniid beetle. Its type species is Pseudobrienia rasnitsyni.PseudoglaesotropisGen. et sp. novValidLegalovEoceneBaltic amberEurope (Baltic Sea region)An anthribid beetle. The type species is Pseudoglaesotropis martynovi. Legalov (2019) subsequently changed the rank of the genus Pseudoglaesotropis to the subgenus of the genus Glaesotropis, and transferred the species P. martynovi to the latter genus.Andrei A. Legalov 2019 A review of the Curculionoidea (Coleoptera) from European Eocene ambers Geosciences 10 1 16 10.3390/geosciences10010016 2019Geosc..10...16L free PulcherhybosorusZhuo Yan Ming Bai Dong Ren 2012 A new fossil Hybosoridae (Coleoptera: Scarabaeoidea) from the Yixian Formation of China Zootaxa 3478 201–204 10.11646/zootaxa.3478.1.21 https://www.mapress.com/zootaxa/2012/f/z03478p204f.pdf Gen. et sp. novValidYan, Bai & RenEarly CretaceousYixian FormationChinaA hybosorid beetle. The type species is Pulcherhybosorus tridentatus.Rhyzobius antiquusSp. novValidKirejtshuk & NelEarliest EoceneOise amberFranceA cocciduline coccinellid beetle, a species of RhyzobiusRhyzobius gratiosusSp. novValidKirejtshuk & NelEarliest EoceneOise amberFranceA cocciduline coccinellid beetle, a species of RhyzobiusSeptiventerMing Bai Rolf G. Beutel Chung-Kun Shih Dong Ren Xing-Ke Yang 2013 Septiventeridae, a new and ancestral fossil family of Scarabaeoidea (Insecta: Coleoptera) from the Late Jurassic to Early Cretaceous Yixian Formation Journal of Systematic Palaeontology 11 3 359–374 10.1080/14772019.2012.660995 2013JSPal..11..359B 84380388 Gen. et sp. novValidBai et al.Late Jurassic or Early CretaceousYixian FormationChinaA septiventerid scarabaeoid beetle. The type species is Septiventer quadridentatus.ShartegopsisA. G. Kirejtshuk 2012 The first record of the family Cleridae (Coleoptera): Shartegopsis miranda gen. et sp. nov. in the mesozoic (Shar-Teg, Upper Jurassic, Southwestern Mongolia) Paleontological Journal 46 6 601–606 10.1134/S0031030112060056 2012PalJ...46..601K 86161456 Gen. et sp. novValidKirejtshukLate JurassicMongoliaA clerid beetle. The type species is Shartegopsis miranda.SinodromeusGen. et sp. novValidWang, Zhang & PonomarenkoEarly CretaceousYixian FormationChinaA trachypachid. The type species is Sinodromeus liutiaogouensis.SinopeltisYali Yu Richard A. B. Leschen Adam Slipinski Dong Ren Hong Pang 2012 The First Fossil Bark-Gnawing Beetle from the Middle Jurassic of Inner Mongolia, China (Coleoptera: Trogossitidae) Annales Zoologici 62 2 245–252 10.3161/000345412X652765 86479829 Gen. et 2 sp. novValidYu et al.Middle JurassicJiulongshan FormationChinaA peltine trogossitid beetle. Genus contains two species: S. jurrasica (type species) and S. amoena. Kirejtshuk (2017) considered the genus Sinopeltis to be a junior synonym of the genus Cervicatinius Tan & Ren (2007).A.G. Kirejtshuk 2017 Taxonomic notes on fossil beetles (Insecta: Coleoptera) Russian Entomological Journal 26 1 35–36 10.15298/rusentj.26.1.03 free SinoschizalaE.A. Jarzembowski E.V. Yan Bo Wang Hai-Chun Zhang 2012 A new flying water beetle (Coleoptera: Schizophoridae) from the Jurassic Daohugou lagerstätte Palaeoworld 21 3–4 160–166 10.1016/j.palwor.2012.09.002Gen. et sp. novValidJarzembowski et al.Middle JurassicDaohugou BedsChinaA schizophorid beetle. The type species is Sinoschizala darani.SuccinometrioxenaAndrei A. Legalov 2012 A new weevil genus of the tribe Metrioxenini (Coleoptera: Belidae) in Eocene Baltic amber Historical Biology: An International Journal of Paleobiology 24 2 213–217 10.1080/08912963.2011.602404 2012HBio...24..213L 84817408 Gen. et sp. novValidLegalovEoceneEuropeA belid weevil found in Baltic amber. The type species is Succinometrioxena poinari.Sucinoptinus bukejsiVitaly I. Alekseev 2012 Sucinoptinus bukejsi sp.nov. (Coleoptera:Ptinidae:Ptinini), the second species of the Tertiary genus from the Baltic amber Baltic Journal of Coleopterology 12 2 145–148 http://www.bjc.sggw.pl/viewart.php?bookid=23&artid=448 Sp. novValidAlekseevEocene (Priabonian)Prussian FormationRussia (Kaliningrad Oblast)A spider beetle found in Baltic amber, a species of Sucinoptinus.TafforeusGen. et sp. novValidPerreauEoceneRussia (Kaliningrad Oblast)A leiodid beetle found in Baltic amber. The type species is Tafforeus cainosternus.TalbragarusRolf G. Oberprieler Stefanie K. Oberprieler 2012 Talbragarus averyi gen. et sp. n., the first Jurassic weevil from the southern hemisphere (Coleoptera: Curculionoidea: Nemonychidae) Zootaxa 3478 256–266 10.11646/zootaxa.3478.1.25 https://www.mapress.com/zootaxa/2012/f/z03478p266f.pdf Gen. et sp. novValidOberprieler & OberprielerLate JurassicTalbragar Fish BedAustraliaA nemonychid weevil. The type species is Talbragarus averyi.Tetraphalerus decorosusSp. novTan et al.Middle JurassicJiulongshan FormationChinaOriginally described as a species of Tetraphalerus; Kirejtshuk (2020) transferred this species to the ommatine genus Monticupes.Alexander G. Kirejtshuk 2020 Taxonomic review of fossil coleopterous families (Insecta, Coleoptera). Suborder Archostemata: Superfamilies Coleopseoidea and Cupedoidea Geosciences 10 2 73 10.3390/geosciences10020073 2020Geosc..10...73K free ToksuniusA. G. Ponomarenko E. V. Yan Bo Wang Hai-chun Zhang 2012 Revision of some early Mesozoic beetles from China Acta Palaeontologica Sinica 51 4 475–490 http://eng.oversea.cnki.net/kcms/detail/detail.aspx?dbCode=cjfd&QueryID=229&CurRec=7&filename=GSWX201204009&dbname=CJFD2012 2016-02-06 2016-06-04 https://web.archive.org/web/20160604044254/http://eng.oversea.cnki.net/kcms/detail/detail.aspx?dbCode=cjfd&QueryID=229&CurRec=7&filename=GSWX201204009&dbname=CJFD2012 dead Gen. et comb. novValidPonomarenko et al.Late TriassicChinaA new genus for "Ademosynoides" chinaticus Lin (1992).Tomoderus balticusDmitry Telnov 2012 Tomoderinae (Coleoptera: Anthicidae) of the Baltic amber Latvijas Entomologs 51 3–11 http://leb.daba.lv/Tomoderinae.pdfSp. novValidTelnovLate EoceneLatvia or LithuaniaA tomoderine anthicid beetle found in Baltic amber, a species of Tomoderus.Trixagus majusculusA.V. Kovalev A.G. Kirejtshuk A. Nel 2012 New species of the genus Trixagus Kugelann, 1794 (Coleoptera: Throscidae) from the lowermost Eocene amber of Oise (France) https://www.zin.ru/journals/trudyzin/doc/vol_316_1/TZ_316_1_Kovalev.pdf Proceedings of the Zoological Institute of the Russian Academy of Sciences 316 1 83–88 sp novValidKovalev, Kirejtshuk & NelLowermost EoceneFranceA throscid beetle, a species of Trixagus.Unda chifengensissp novValidWang, Zhang & PonomarenkoMiddle JurassicDaohugou BedsChinaA trachypachid, a species of Unda.XenohimatiumG.Yu. Lyubarsky E.E. Perkovsky 2012 A new genus of Erotylidae from Eocene amber (Coleoptera: Clavicornia) Russian Entomological Journal 21 1 35–38 10.15298/rusentj.21.1.04 free Gen. et sp. novValidLyubarsky & PerkovskyLate EoceneRovno amberUkraineA pleasing fungus beetle belonging to the subfamily Xenoscelinae. The type species is X. rovnense. Diptera Name Novelty Status Authors Age Unit Location Notes ImagesAetheapnomyiaGen. et comb. novValidHarbach & GreenwaltEoceneEuropeA mosquito found in Baltic Amber, a new genus for "Aedes" hoffeinsorum Szadziewski 1998.Bibio anatolicusJohn Skartveit André Nel 2012 Fossil Bibionidae (Diptera: Bibionomorpha) from the late Oligocene of Bes-Konak, Anatolia, Turkey Zootaxa 3329 1 51–63 10.11646/zootaxa.3329.1.5 Sp. novValidSkartveit & NelOligoceneTurkeyA species of Bibio.Bibio nigricostaSp. novValidSkartveit & NelOligoceneTurkeyA species of Bibio.Calosargus talbragarensisStefanie K. Oberprieler David K. Yeates 2012 Calosargus talbragarensis New Species: the First Brachyceran Fly from the Jurassic of Australia (Diptera, Archisargidae) Journal of Paleontology 86 4 641–645 http://jpaleontol.geoscienceworld.org/content/86/4/641.abstract 10.1666/11-126R.1 2012JPal...86..641O 128737218 subscription Sp. nov ValidOberprieler & YeatesLate JurassicAustraliaAn archisargid brachyceran, a species of Calosargus.ClarumredderaZ. A. Fedotova E. E. Perkovsky 2012 Gall midges of the supertribe Stomatosematidi (Diptera, Cecidomyiidae) in the Palaearctic Region, with descriptions of new taxa from the Late Eocene Rovno amber: 1. Stomatosema, Vanchidiplosis, Clarumreddera gen. n. Entomological Review 92 4 427–439 10.1134/S0013873812040082 2012EntRv..92..427F 84879200 Gen. et 2 sp. novValidFedotova & PerkovskyLate EoceneRovno amberUkraineA cecidomyiid fly, with two species: C. korneyevi and C. conceptiva.Culiseta kishenehnRalph E. Harbach Dale Greenwalt 2012 Two Eocene species of Culiseta (Diptera: Culicidae) from the Kishenehn Formation in Montana Zootaxa 3530 25–34 https://www.mapress.com/zootaxa/2012/f/zt03530p034.pdf 10.11646/zootaxa.3530.1.2 free Sp. novValidHarbach & GreenwaltEoceneKishenehn FormationUnited StatesA mosquito, a species of Culiseta.Culiseta lemniscataSp. novValidHarbach & GreenwaltEoceneKishenehn FormationUnited StatesA mosquito, a species of Culiseta.Curtonotum electrodominicumDavid A. Grimaldi Ashley H. Kirk-Spriggs 2012 Fossil Curtonotidae (Diptera: Schizophora: Ephydroidea) American Museum Novitates 3760 1–16 10.1206/3760.2 2246/6392 56131379 https://www.biodiversitylibrary.org/bibliography/166183 Sp. novValidGrimaldi & Kirk-SpriggsMioceneDominican RepublicA curtonotid found in Dominican amber, a species of Curtonotum.DaohugosargusJunfeng Zhang 2012 Orientisargidae fam. n., a new Jurassic family of Archisargoidea (Diptera, Brachycera), with review of Archisargidae from China ZooKeys 238 57–76 10.3897/zookeys.238.3624 23226708 3496945 free 2012ZooK..238...57J Gen. et comb. novValidZhangMiddle JurassicDaohugou BedsChinaAn archisargid brachyceran; a new genus for "Sharasargus" eximius Zhang, Yang & Ren (2008).DepressonotumGen. et sp. novValidGrimaldi & Kirk-SpriggsMioceneDominican RepublicA curtonotid found in Dominican amber. The type species is Depressonotum priscum.Dicranomyia (Caenolimonia) alexbrowniSigitas Podenas George O. Poinar, Jr. 2012 New Short-Palped Crane Flies (Diptera: Limoniidae) from Mexican Amber Proceedings of the Entomological Society of Washington 114 3 347–371 10.4289/0013-8797.114.3.347 85216822 Sp. novValidPodenas & PoinarLate Oligocene to early MioceneMexicoA species of Dicranomyia.Dicranomyia (Dicranomyia) chiapaSp. novValidPodenas & PoinarLate Oligocene to early MioceneMexicoA species of Dicranomyia.Dicranomyia (Dicranomyia) mexaSp. novValidPodenas & PoinarLate Oligocene to early MioceneMexicoA species of Dicranomyia.Dicranomyia (Dicranomyia) vellaSp. novValidPodenas & PoinarLate Oligocene to early MioceneMexicoA species of Dicranomyia.Didactylomyia dlusskyiZ. A. Fedotova E. E. Perkovsky 2012 Gall midges of the supertribe Stomatosematidi (Diptera, Cecidomyiidae) in the Palaearctic region, with descriptions of new taxa from the late Eocene Rovno Amber. 2. The genera Didactylomyia and Rovnodidactylomyia gen. n. Entomological Review 92 5 565–575 10.1134/S0013873812050090 2012EntRv..92..565F 6376732 Sp. novValidFedotova & PerkovskyLate EoceneUkraineA cecidomyiid fly found in Rovno amber, a species of Didactylomyia.EnischnomyiaPoinar G. O. Jr. Brown A. 2012 The first fossil streblid bat fly, Enischnomyia stegosoma n. g., n. sp. (Diptera: Hippoboscoidea: Streblidae) Systematic Parasitology 81 2 79–86 10.1007/s11230-011-9339-2 22183917 14469619 Gen. et sp. novValidPoinar & BrownBurdigalianDominican amberDominican RepublicA bat fly, type species E. stegosoma, host for the Plasmodiidae Vetufebrus ovatusPoinar G. O. Jr. 2011 Vetufebrus ovatus n. gen., n. sp. (Haemospororida: Plasmodiidae) vectored by a streblid bat fly (Diptera: Streblidae) in Dominican amber Parasites & Vectors 4 229 229 10.1186/1756-3305-4-229 22152687 3253689 free thumb|center|150px|Enischnomyia stegosomaEoaedesGen. et comb. novValidHarbach & GreenwaltEoceneEuropeA mosquito found in Baltic Amber, a new genus for "Aedes" damzeni Szadziewski 1998.Eoptychopterina posticaLuxi Liu Chungkun Shih Dong Ren 2012 Two new species of Ptychopteridae and Trichoceridae from the Middle Jurassic of northeastern China (Insecta: Diptera: Nematocera) Zootaxa 3501 55–62 https://www.mapress.com/zootaxa/2012/f/z03501p062f.pdf 10.11646/zootaxa.3501.1.2 Sp. novDisputedLiu, Shih & RenMiddle JurassicJiulongshan FormationChinaA ptychopterid, a species of Eoptychopterina. Considered to be a junior synonym of Eoptychopterina elenae Ren & Krzemiński (2002) by Lukashevich & Ansorge (2026).Elena D. Lukashevich Jörg Ansorge First Eoptychopterina (Diptera: Ptychopteridae) from the Jurassic of Europe, with taxonomic notes on Eoptychopterininae and Eoptychopterinae 2026 Palaeoentomology 9 5 440–449 10.11646/palaeoentomology.9.5.12 Eotrichocera (Archaeotrichocera) spatiosaSp. novValidLiu, Shih & RenMiddle JurassicJiulongshan FormationChinaA trichocerid, a species of Eotrichocera.Erioptera (?Mesocyphona) beataSp. novValidPodenas & PoinarLate Oligocene to early MioceneMexicoA species of Erioptera.Erioptera (Mesocyphona) diverSp. novValidPodenas & PoinarLate Oligocene to early MioceneMexicoA species of Erioptera.Erioptera (?Mesocyphona) sherrySp. novValidPodenas & PoinarLate Oligocene to early MioceneMexicoA species of Erioptera.FlagellisargusJunfeng Zhang 2012 Distinct But Rare Archisargid Flies from Jurassic of China (Diptera, Brachycera, Archisargidae) With Discussion of the Systematic Position of Origoasilus pingquanensis ZHANG ET AL., 2011 Journal of Paleontology 86 5 878–885 10.1666/12-028.1 2012JPal...86..878Z 86217269 Gen. et 3 sp. novValidZhangCallovian or OxfordianDaohugou BedsChinaAn archisargid fly. Genus contains three species: Flagellisargus sinicus, F. venustus and F. robustus.GroveromyiaZ. A. Fedotova E. E. Perkovsky 2012 Gall midges of the supertribe Stomatosematidi (Diptera, Cecidomyiidae) in the Palaearctic region, with description of new taxa from the Late Eocene Rovno amber: 3. Groveromyia gen. n. analysis of biometric parameters of the recent and late Eocene genera and species Entomological Review 92 8 932–948 10.1134/S001387381208012X 2012EntRv..92..932F 17772683 Gen. et 3 sp. et comb. novValidFedotova & PerkovskyLate EoceneRovno amberUkraineA cecidomyiid fly with 4 species G. occlusa; G. digna, G. astrosa, and "Bryocrypta" concinna Fedotova (2005).Helius (Helius) collemusSp. novValidPodenas & PoinarLate Oligocene to early MioceneMexicoA species of Helius.Jurochlus adustusElena D. Lukashevich Andrey A. Przhiboro 2012 Pupae of Mesozoic Jurochlus Kalugina, 1985 (Diptera: Chironomidae), with description of four new species Zootaxa 3478 434–452 10.11646/zootaxa.3478.1.39 https://www.mapress.com/zootaxa/2012/f/z03478p452f.pdf Sp. novValidLukashevich & PrzhiboroMesozoicMongoliaA chironomid, a species of Jurochlus.Jurochlus limbatusSp. novValidLukashevich & PrzhiboroMesozoicMongoliaA chironomid, a species of Jurochlus.Jurochlus lineatusSp. novValidLukashevich & PrzhiboroMesozoicMongoliaA chironomid, a species of Jurochlus.Jurochlus trivittatusSp. novValidLukashevich & PrzhiboroMesozoicMongoliaA chironomid, a species of Jurochlus.LacrimyzaJindřich Roháček 2013 New amber fossil Anthomyzidae (Diptera): an unexpected Eocene diversity Journal of Systematic Palaeontology 11 4 431–473 10.1080/14772019.2012.691907 2013JSPal..11..431R 84457598 Gen. et 2 sp. novValidRoháčekEoceneEuropeAn anthomyzid. Genus contains two species: L. lacrimosa and L. christelae.LaiyangitabanusJunfeng Zhang 2012 New horseflies and water snipe-flies (Diptera: Tabanidae and Athericidae) from the Lower Cretaceous of China Cretaceous Research 36 1–5 10.1016/j.cretres.2012.01.004 2012CrRes..36....1Z Gen. et sp. novValidZhangEarly CretaceousLaiyang FormationChinaA horse-fly. The type species is Laiyangitabanus formosus.LithorhagioJ. -f. Zhang H. -j. Li 2012 New taxa of snipe flies (Diptera: Brachycera: Rhagionidae) in the Daohugou Biota, China Paleontological Journal 46 2 157–163 10.1134/S0031030112020128 2012PalJ...46..157Z 84452309 Gen. et sp. novValidZhang & LiMiddle Jurassic or lowermost Upper Jurassic Daohugou BedsChinaA rhagionid. The type species is Lithorhagio megalocephalus.Lygistorrhina caribbianaMartin Grund 2012 A Long-Beaked Fungus Gnat in Dominican Amber (Diptera: Lygistorrhinidae) Annales Zoologici 62 4 639–642 10.3161/000345412X659704 85886454 Sp. novValidGrundMioceneDominican RepublicA lygistorrhinid fungus gnat found in Dominican amber, a species of Lygistorrhina.Mesorhyphus handlirschiE. D. Lukashevich 2012 New Bibionomorpha (Insecta: Diptera) from the Jurassic of Asia Paleontological Journal 46 3 273–287 10.1134/S0031030112030100 2012PalJ...46..273L 86768109 Sp. novValidLukashevichUpper Jurassic MongoliaAn anisopodid bibionomorph, a species of Mesorhyphus.Mesorhyphus hennigiSp. novValidLukashevichUpper Jurassic MongoliaAn anisopodid bibionomorph, a species of Mesorhyphus.Mesosciophila sigmoideaQian Wang Yunyun Zhao Dong Ren 2012 Two new species of Mesosciophilidae (Insecta: Diptera: Nematocera) from the Yanliao biota of Inner Mongolia, China Alcheringa: An Australasian Journal of Palaeontology 36 4 509–514 10.1080/03115518.2012.685434 2012Alch...36..509W 128823650 sp novValidWang, Zhao & RenMiddle Jurassic Jiulongshan FormationChinaA mesosciophilid nematoceran, a species of Mesosciophila.Mesosolva zhangaeNom. novValidZhangMiddle JurassicDaohugou BedsChinaAn archisargid brachyceran; a replacement name for Brevisolva daohugouensis Zhang et al. (2010).Nemopalpus inexpectatusRüdiger Wagner Brian R. Stuckenberg 2012 New fossil and extant species of Nemopalpus Macquart (Diptera: Psychodidae, Bruchomyiinae) African Invertebrates 53 1 355–367 10.5733/afin.053.0119 free 2012AfrIn..53..355W Sp. novValidWagner in Wagner & StuckenbergLate EoceneBaltic amberEuropeA bruchomyiine psychodid fly. Originally described as a species of Nemopalpus, but subsequently transferred to the genus Palaeosycorax.Rüdiger Wagner 2017 Synopsis of extinct Bruchomyiinae (Diptera, Psychodidae) from Burmese, Baltic and Dominican amber, with descriptions of new genera and species Zootaxa 4320 1 100–120 10.11646/zootaxa.4320.1.6 Nemopalpus velteniSp. novValidWagner in Wagner & StuckenbergCretaceous (Albian)Burmese amberMyanmarA bruchomyiine psychodid fly. Originally described as a species of Nemopalpus, but subsequently transferred to the genus Palaeoglaesum.OrientisargusGen. et sp. novValidZhangLatest Middle Jurassic or earliest Late JurassicDaohugou BedsChinaAn orientisargid archisargoid brachyceran. The type species is Orientisargus illecebrosus.Oryctochlus brundiniElena D. Lukashevich 2012 Pupae of Mesozoic Oryctochlus Kalugina, 1985 (Chironomidae: Podonominae), with description of two new species Fauna Norvegica 31 159–165 10.5324/fn.v31i0.1400 free Sp. novValidLukashevichLate JurassicMongoliaA podonomine chironomid, as species of OryctochlusOryctochlus kaluginaeSp. novValidLukashevichLate JurassicMongoliaA podonomine chironomid, as species of OryctochlusPalaeopoecilostola eocenicaIwona Kania Wiesław Krzemiński 2012 A new species of Palaeopoecilostola Meunier, 1899 (Diptera: Limoniidae) from the Eocene Baltic amber Zootaxa 3495 42–56 https://www.mapress.com/zootaxa/2012/f/z03495p056f.pdf 10.11646/zootaxa.3495.1.2 Sp. novValidKania & KrzemińskiEoceneEuropeA limoniid found in Baltic amber, a species of Palaeopoecilostola.Paramesosciophilodes aequussp novValidWang, Zhao & RenMiddle Jurassic Jiulongshan FormationChinaA mesosciophilid nematoceran, a species of Paramesosciophilodes.Penthetria beskonakensisSp. novValidSkartveit & NelOligoceneTurkeyA species of Penthetria.Probittacomorpha brisaciWiesław Krzemiński Iwona Kania André Nel 2012 Probittacomorpha brisaci n. sp. (Ptychopteridae, Bittacomorphinae) from the Late Miocene of Montagne d'Andance, Saint Bauzile, Ardèche (France) Zootaxa 3521 80–88 https://www.mapress.com/zootaxa/2012/f/z03521p088f.pdf 10.11646/zootaxa.3521.1.6 sp novValidKrzemiński, Kania & NelLate MioceneFranceA ptychopterid, a species of Probittacomorpha.Procramptonomyia kovaleviSp. novValidLukashevichUpper Jurassic Kazakhstan MongoliaA procramptonomyiid bibionomorph, a species of Procramptonomyia.Procramptonomyia ponomarenkoiSp. novValidLukashevichUpper Jurassic MongoliaA procramptonomyiid bibionomorph, a species of Procramptonomyia.Protanthomyza hennigiSp. novValidRoháčekEoceneEuropeAn anthomyzid, a species of Protanthomyza.Protanthomyza hoffeinsorumSp. novValidRoháčekEoceneEuropeAn anthomyzid, a species of Protanthomyza.Protanthomyza kryloviSp. novValidRoháčekEoceneEuropeAn anthomyzid, a species of Protanthomyza.Protanthomyza loewiSp. novValidRoháčekEoceneEuropeAn anthomyzid, a species of Protanthomyza.Protanthomyza meunieriSp. novValidRoháčekEoceneEuropeAn anthomyzid, a species of Protanthomyza.Protanthomyza presliSp. novValidRoháčekEoceneEuropeAn anthomyzid, a species of Protanthomyza.Protanthomyza tschirnhausiSp. novValidRoháčekEoceneEuropeAn anthomyzid, a species of Protanthomyza.Protoculicoides ciliatusArt Borkent 2012 Further biting midges (Diptera: Ceratopogonidae) in Canadian Cretaceous amber The Canadian Entomologist 144 6 834–842 10.4039/tce.2012.83 85002252 Sp. nov ValidBorkentCretaceousCanadian amberCanadaA member of the family Ceratopogonidae. Originally described as a species of Protoculicoides; Borkent (2019) transferred this species to the genus Atriculicoides.Art Borkent 2019 The phylogenetic relationships of Cretaceous biting midges, with a key to all known genera (Diptera, Ceratopogonidae) American Museum Novitates 3921 1–48 10.1206/3921.1 2246/6927 91737913 https://www.biodiversitylibrary.org/item/266374 Protorhagio parvussp novValidZhang & LiMiddle Jurassic or lowermost Upper Jurassic Daohugou BedsChinaA rhagionid, a species of Protorhagio.Protorhyphus rohdendorfiSp. novValidLukashevichUpper Jurassic MongoliaA protorhyphid bibionomorph, a species of Protorhyphus.RovnodidactylomyiaGen. et 3 sp. et comb. novValidFedotova & PerkovskyLate EoceneEuropeA cecidomyiid fly with 5 species R. zosimovichi, R. sidorenkoi, R. iconica) in Rovno amber, plus "Bryocrypta" girafa Meunier (1904) and "B." capitosa Meunier (1904) in Baltic amber.SinallomyiaNom. novValidZhangEarly CretaceousYixian FormationChinaAn archisargid brachyceran; a replacement name for Allomyia Ren (1998).SinocretomyiaGen. et sp. novValidZhangEarly CretaceousLaiyang FormationChinaAn athericid. The type species is Sinocretomyia minuscula.Stilobezzia pikeiSp. nov ValidBorkentCretaceousCanadian amberCanadaA species of Stilobezzia.Stomatosema iljieteugeniaeSp. novValidFedotova & PerkovskyLate EoceneRovno amberUkraineA cecidomyiid flyTipula paleopannoniaMichael S. Engel Martin Gross 2012 A New Fossil Crane Fly from the Early Pannonian of the Styrian Basin (Diptera: Tipulidae) Journal of the Kansas Entomological Society 85 2 160–163 10.2317/JKES120412.1 86013911 Sp. novValidEngel & GrossMiocene (ca. 11.3 Ma)Paldau FormationAustriaA crane fly, a species of Tipula. Ephemeroptera Name Novelty Status Authors Age Unit Location Notes ImagesAlexandriniaSinitshenkova N. D. Vassilenko D. V. 2012 The latest record of mayflies of the family Protereismatidae Sellards (Ephemerida = Ephemeroptera) and a new species of the family Misthodotidae in the Upper Permian of Europe Paleontological Journal 46 1 61–65 10.1134/S0031030112010121 2012PalJ...46...61S 86776832 Gen. et sp.ValidSinitshenkova & VassilenkoPermian Poldarsa FormationRussia United StatesA mayfly. Species are A. gigantea and A. directum.HammephemeraAlexei Bashkuev Jürgen Sell Daniil Aristov Alexandr Ponomarenko Nina Sinitshenkova Horst Mahler 2012 Insects from the Buntsandstein of Lower Franconia and Thuringia Paläontologische Zeitschrift 86 2 175–185 10.1007/s12542-011-0119-8 2012PalZ...86..175B 55603192 Gen. et sp. nov ValidSinitshenkova in Bashkuev et al.Early TriassicGermanyA sharephemerid mayfly. The type species is Hammephemera pulchra.Misthodotes tshernovaesp novValidSinitshenkova & VassilenkoUpper Permian Poldarsa FormationRussiaA mayfly, a species of Misthodotes.PonalexN. D. Sinitshenkova D. S. Aristov 2012 The biggest fossil mayfly (Insecta: Ephemerida = Ephemeroptera) from the Upper Permian locality of Isady, northern European Russia Far Eastern Entomologist 255 8–10 http://www.biosoil.ru/fee/2012/N-255/N-255.pdf 2012-12-20 2013-10-20 https://web.archive.org/web/20131020033339/http://www.biosoil.ru/fee/2012/N-255/N-255.pdf dead Gen. et sp. novValidSinitshenkova & AristovLate PermianPoldarsa FormationRussiaA protereismatid mayfly. The type species is Ponalex maximus.Siphloplecton barabaniArnold H. Staniczek Roman J. Godunko 2012 Revision of fossil Metretopodidae (Insecta: Ephemeroptera) in Baltic amber – Part 1: Type specimens of hitherto described fossil species of Siphloplecton CLEMENS, 1915 and Metretopus EATON, 1901; with description of four new fossil species of Siphloplecton Palaeodiversity 5 57–87 https://www.palaeodiversity.org/pdf/05/Palaeodiversity_5_57-87_Stanicek.pdfSp. novValidStaniczek & GodunkoEoceneEuropeA metretopodid mayfly found in Baltic amber, a species of Siphloplecton.Siphloplecton demouliniSp. novValidStaniczek & GodunkoEoceneEuropeA metretopodid mayfly found in Baltic amber, a species of Siphloplecton.Siphloplecton hageniSp. novValidStaniczek & GodunkoEoceneEuropeA metretopodid mayfly found in Baltic amber, a species of Siphloplecton.Siphloplecton pictetiSp. novValidStaniczek & GodunkoEoceneEuropeA metretopodid mayfly found in Baltic amber, a species of Siphloplecton. Hemiptera Name Novelty Status Authors Age Unit Location Notes ImagesAcaricoris robertaeErnst Heiss George O. Poinar Jr. 2012 The first Carventinae species in Miocene Dominican Amber (Hemiptera: Heteroptera: Aradidae) Zootaxa 3268 47–54 https://www.mapress.com/zootaxa/2012/f/z03268p054f.pdf 10.11646/zootaxa.3268.1.5 Sp. novValidHeiss & PoinarMioceneDominican RepublicAn aradid found in Dominican amber, a species of Acaricoris.Arctorthezia balticaIsabelle M. Vea David A. Grimaldi 2012 Phylogeny of ensign scale insects (Hemiptera: Coccoidea: Ortheziidae) based on the morphology of recent and fossil females Systematic Entomology 37 4 758–783 10.1111/j.1365-3113.2012.00638.x 2012SysEn..37..758V 84368560 Sp. novValidVea & GrimaldiEoceneEuropeAn ortheziid found in Eocene Baltic amber, a species of Arctorthezia.BreviscutumYunzhi Yao Wanzhi Cai David A. Rider Dong Ren 2013 Primipentatomidae fam. nov. (Hemiptera: Heteroptera: Pentatomomorpha), an extinct insect family from the Cretaceous of north-eastern China Journal of Systematic Palaeontology 11 1 63–82 10.1080/14772019.2011.639814 2013JSPal..11...63Y 86753502 Gen. et sp. novValidYao et al.CretaceousChinaA primipentatomid pentatomoid. The type species is Breviscutum lunatum.BurmacicadaGeorge Poinar Jr. Gene Kritsky 2012 Morphological conservatism in the foreleg structure of cicada hatchlings, Burmacicada protera n. gen., n. sp. in Burmese amber, Dominicicada youngi n. gen., n. sp. in Dominican amber and the extant Magicicada septendecim (L.) (Hemiptera: Cicadidae) Historical Biology: An International Journal of Paleobiology 24 5 461–466 10.1080/08912963.2011.603421 2012HBio...24..461P 82426789 Gen. et sp. novValidPoinar & KritskyCretaceousBurmese amberMyanmarA cicada. The type species is Burmacicada protera.BurmortheziaGen et 2 sp. novValidVea & GrimaldiCretaceous (98 Ma)Burmese amberMyanmarAn ortheziid. Genus contains two species: Burmorthezia kotejai and B. insolita.Carventus bechlyiSp. novValidHeiss & PoinarMioceneDominican RepublicAn aradid found in Dominican amber, a species of Carventus.Cicadocoris variansQiuping Dong Yunzhi Yao Dong Ren 2012 A new species of Progonocimicidae (Hemiptera: Coleorrhyncha) from northeastern China Zootaxa 3495 73–78 https://mapress.com/zootaxa/2012/f/z03495p078f.pdf 10.11646/zootaxa.3495.1.4 Sp. novDisputedDong, Yao & RenMiddle JurassicJiulongshan FormationChinaA moss bug belonging to the family Progonocimicidae. Jiang & Huang (2017) considered this species to be a probable junior synonym of Cicadocoris sinensis Hong (1983).Jia-Qian Jiang Di-Ying Huang 2017 New species of Cicadocoris (Hemiptera: Coleorrhyncha: Progonocimicidae) from mid-Jurassic deposits in northeastern China European Journal of Entomology 114 355–364 10.14411/eje.2017.045 free CinemalaDmitry E. Shcherbakov 2012 A new subfamily of Mesozoic Hylicellidae (Homoptera: Cicadomorpha) Russian Entomological Journal 21 4 441–444 10.15298/rusentj.21.4.10 free Gen. et sp. novValidShcherbakovMiddle or Late JurassicBakhar GroupMongoliaA member of Cicadomorpha belonging to the family Hylicellidae. The type species is C. contrasta.CintuxStroiński Adam Szwedo Jacek 2012 The oldest known Lophopidae planthopper (Hemiptera: Fulgoromorpha) from the European Palaeocene Geobios 45 4 413–420 10.1016/j.geobios.2011.10.007 2012Geobi..45..413S Gen. et sp. novValidStroiński & SzwedoPalaeoceneFranceA lophopid planthopper. The type species is Cintux menatensis.ConjucellaGen. et sp. novValidShcherbakovEarly CretaceousZaza FormationRussiaA member of Cicadomorpha belonging to the family Hylicellidae. The type species is C. selene.DellasharaDmitry E. Shcherbakov 2012 More on Mesozoic Membracoidea (Homoptera) Russian Entomological Journal 21 1 15–22 10.15298/rusentj.21.1.02 free Gen. et sp. novValidShcherbakovLate JurassicShar Teg SequenceMongoliaA member of Cicadomorpha belonging to the superfamily Membracoidea and the family Archijassidae. The type species is D. tega.DominicicadaGen. et sp. novValidPoinar & KritskyDominican RepublicA cicada found in Dominican amber. The type species is Dominicicada youngi.Edessa proteraGeorge Poinar Jr. Donald B. Thomas 2012 A stink bug, Edessa protera sp. n. (Pentatomidae: Edessinae) in Mexican amber Historical Biology: An International Journal of Paleobiology 24 2 207–211 10.1080/08912963.2011.602403 2012HBio...24..207P 84081033 Sp. novValidPoinar & ThomasTertiaryMexicoA pentatomid stink bug found in Mexican amber, a species of Edessa.HispanocaderV.B. Golub Yu.A. Popov A. Arillo 2012 Hispanocaderidae n. fam. (Hemiptera: Heteroptera: Tingoidea), one of the oldest lace bugs from the Lower Cretaceous Álava amber (Spain) Zootaxa 3270 41–50 https://www.mapress.com/zootaxa/2012/f/z03270p050f.pdf 10.11646/zootaxa.3270.1.3 Gen. et sp. novValidGolub, Popov & ArilloAlbianEscucha FormationSpainA relative of tingids. The type species is Hispanocader lisae.HoffeinsoriaAleksander Herczek Yuri A. Popov 2012 A new peculiar isometopine genus (Hemiptera: Heteroptera: Miridae) from the Eocene Baltic amber Zootaxa 3196 64–68 https://www.mapress.com/zootaxa/2012/f/z03196p068f.pdf 10.11646/zootaxa.3196.1.4 Gen. et sp. novValidHerczek & PopovEocenePrussian FormationEuropeAn isometopine mirid found in Eocene Baltic amber. The type species is Hoffeinsoria robusta.JianiaBo Wang Jacek Szwedo Haichun Zhang 2012 New Jurassic Cercopoidea from China and their evolutionary significance (Insecta: Hemiptera) Palaeontology 55 6 1223–1243 10.1111/j.1475-4983.2012.01185.x 2012Palgy..55.1223W 86793512 free Gen. et 2 sp. novValidWang & Szwedo in Wang, Szwedo & ZhangMiddle JurassicDaohugou BedsChinaA sinoalid froghopper. Genus contains two species: Jiania crebra and Jiania gracila. Fu, Cai & Huang (2018) considered J. gracila to be a junior synonym of J crebra.Yanzhe Fu Chenyang Cai Diying Huang 2018 A new fossil sinoalid species from the Middle Jurassic Daohugou beds (Insecta: Hemiptera: Cercopoidea) Alcheringa: An Australasian Journal of Palaeontology 42 1 94–100 10.1080/03115518.2017.1374458 2018Alch...42...94F 135402986 KachinocorisErnst Heiss 2012 Kachinocoris brevipennis n.gen., n.sp. in Cretaceous Burmese Amber (Hemiptera: Heteroptera: Aradidae) Zootaxa 3227 64–68 https://www.mapress.com/zootaxa/2012/f/z03227p068f.pdf 10.11646/zootaxa.3227.1.4 Gen. et sp. novValidHeissCretaceousBurmese amberMyanmarA flat bug found in Cretaceous Burmese Amber. The type species is Kachinocoris brevipennis.KemobiusGen. et sp. novValidShcherbakovMiddle JurassicItat FormationRussiaA member of Cicadomorpha belonging to the superfamily Membracoidea and the family Archijassidae. The type species is K. lux.KisaGen. et sp. novValidShcherbakovMiddle JurassicItat FormationRussiaA member of Cicadomorpha belonging to the superfamily Membracoidea and the family Archijassidae. The type species is K. fasciata.KubecolaGen. et sp. novValidShcherbakovMiddle JurassicItat FormationRussiaA member of Cicadomorpha belonging to the superfamily Membracoidea and the family Archijassidae. The type species is K. guttatus.MicroaradusE. Heiss G.O. Poinar 2012 New Aradidae in Mesozoic Burmese Amber (Hemiptera, Heteroptera) Annalen des Naturhistorischen Museums in Wien, Serie A 114 307–316 http://www.nhm-wien.ac.at/verlag/wissenschaftliche_publikationen/annalen_serie_a/4_22 41702114 Gen. et sp. nov ValidHeiss & PoinarMesozoicBurmese amberMyanmarAn archaearadine aradid found in Mesozoic Burmese amber. The type species is Microaradus anticus.MinyscapheusGeorge Poinar, Jr. Gene Kritsky Alex Brown 2012 Minyscapheus dominicanus n. gen., n. sp. (Hemiptera: Cicadidae), a fossil cicada in Dominican amber Historical Biology 24 3 329–333 10.1080/08912963.2011.635791 84254904 Gen. et sp. nov. ValidPoinar, Kritsky & BrownDominican RepublicA cicada found in Dominican amber. The type species is Minyscapheus dominicanus.Mixorthezia dominicanaSp. novValidVea & GrimaldiMioceneDominican RepublicAn ortheziid found in Miocene Dominican amber, a species of Mixorthezia.Mixorthezia kozariSp. novValidVea & GrimaldiMioceneDominican RepublicAn ortheziid found in Miocene Dominican amber, a species of Mixorthezia.MongolocorisO. V. Ryzhkova 2012 New Saldoid bugs of the family Enicocoridae (Hemiptera: Heteroptera: Leptopodomorpha) from the Lower Cretaceous of Mongolia Paleontological Journal 46 5 485–494 10.1134/S0031030112050085 2012PalJ...46..485R 83523509 Gen. et comb. novValidRyzhkovaEarly CretaceousMongoliaAn enicocorid leptopodomorph heteropteran, a new genus for "Enicocoris" tibialis Popov, 1986.MyanmeziraGen. et sp. nov ValidHeiss & PoinarMesozoicBurmese amberMyanmarA mezirine aradid found in Mesozoic Burmese amber. The type species is Myanmezira longicornis.Nesoproxius latocanusSp. novValidHeiss & PoinarMioceneDominican RepublicAn aradid found in Dominican amber, a species of Nesoproxius.Notonecta vetulaWei-Ting Zhang Yun-Zhi Yao Dong Ren 2012 Phylogenetic analysis of a new fossil Notonectidae (Heteroptera: Nepomorpha) from the Late Jurassic of China Alcheringa: An Australasian Journal of Palaeontology 36 2 239–250 10.1080/03115518.2012.628165 2012Alch...36..239Z 129085255 sp novValidZhang, Yao & RenLate Jurassic Chijinqiao FormationChinaA notonectid,a species of Notonecta.OropentatomaGen. et sp. novValidYao et al.CretaceousChinaA primipentatomid pentatomoid. The type species is Oropentatoma epichara.Poljanka hirsutaGuang Yang Yunzhi Yao Dong Ren 2012 A new species of Protopsyllidiidae (Hemiptera, Sternorrhyncha) from the Middle Jurassic of China Zootaxa 3274 36–42 https://www.mapress.com/zootaxa/2012/f/z03274p042f.pdf 10.11646/zootaxa.3274.1.4 sp novValidYang, Yao & RenMiddle Jurassic Jiulongshan FormationChinaA member of Sternorrhyncha, a protopsyllidiid. Originally described as a species of Poljanka, but subsequently transferred to the genus Cicadellopsis.Marina Hakim Minmin Xu Diying Huang 2026 Protopsyllidiidae from the Jiyuan Basin (Henan, China): Taxonomic Updates and Their Biostratigraphic Implications Insects 17 8 824 10.3390/insects17080824 freePrimipentatomaGen. et 2 sp. novValidYao et al.CretaceousChinaA primipentatomid pentatomoid. Genus contains two species: Primipentatoma peregrina and Primipentatoma fangi.ProneoproxiusGen. et sp. novValidHeiss & PoinarMioceneDominican RepublicAn aradid found in Dominican amber. The type species is Proneoproxius cornutus.PurbecellusGen. et comb. novValidShcherbakovEarly CretaceousPurbeck GroupUnited KingdomA member of Cicadomorpha belonging to the superfamily Membracoidea and the family Archijassidae. The type species is "Cicadellium" psocus Westwood (1854).QuadrocorisGen. et sp. novValidYao et al.CretaceousChinaA primipentatomid pentatomoid. The type species is Quadrocoris radius.Sinaldocader rasnitsyniV. B. Golub Yu. A. Popov 2012 A new lace bug species of the genus Sinaldocader (Hemiptera: Heteroptera, Tingidae) from the Turonian of southwestern Kazakhstan Paleontological Journal 46 3 258–261 10.1134/S0031030112020050 2012PalJ...46..258G 86293197 Sp. novValidGolub & PopovTuronian KazakhstanA lace bug. Originally described as a species of Sinaldocader; Popov & Golub (2019) transferred this species to the separate genus Kzylcader.Yu. A. Popov V.B. Golub 2019 A new tribe, genus and four species of lace bugs (Hemiptera: Heteroptera: Tingidae) from the Upper Cretaceous of southwestern Kazakhstan Paleontological Journal 53 2 155–164 10.1134/S0031030119010088 2019PalJ...53..155P 195319420 https://elibrary.ru/item.asp?id=37068675 subscription SinoalaGen. et sp. novValidWang & Szwedo in Wang, Szwedo & ZhangMiddle JurassicDaohugou BedsChinaA sinoalid froghopper. The type species is Sinoala parallelivena.Sunotettigarcta hirsutaShu Li Ying Wang Dong Ren Hong Pang 2012 Revision of the genus Sunotettigarcta Hong, 1983 (Hemiptera, Tettigarctidae), with a new species from Daohugou, Inner Mongolia, China Alcheringa: An Australasian Journal of Palaeontology 36 4 501–507 10.1080/03115518.2012.680722 2012Alch...36..501L 128491887 sp novValidLi, Wang & Ren in Li et al.Middle Jurassic Daohugou BedsChinaA tettigarctid, a species of Sunotettigarcta.UlanocorisGen. et 2 sp. novValidRyzhkovaEarly CretaceousMongoliaAn enicocorid leptopodomorph heteropteran. Genus contains two species: Ulanocoris femoralis and U. grandis.VenustsaldaWeiting Zhang Jingjing Song Yunzhi Yao Dong Ren 2012 A new fossil Saldidae (Hemiptera: Heteroptera: Leptopodomorpha) from the Early Cretaceous in China Zootaxa 3273 63–68 https://www.mapress.com/zootaxa/2012/f/z03273p068f.pdf 10.11646/zootaxa.3273.1.4 Gen. et sp. novValidZhang et al.Lower CretaceousYixian FormationChinaA saldid heteropteran. The type species is Venustsalda locella. Hymenoptera Name Novelty Status Authors Age Unit Location Notes ImagesAgapiaKopylov D. S. 2012 New Ichneumonidae (Hymenoptera) from the Upper Cretaceous ambers of the Taimyr Peninsula Paleontological Journal 46 4 383–391 10.1134/S0031030112040041 2012PalJ...46..383K 85207197 Gen. et sp. novValidKopylovLate CretaceousRussiaAn ichneumonid. The type species is Agapia sukatchevae.AgapteronGen. et sp. novValidKopylovLate CretaceousRussiaAn ichneumonid. The type species is Agapteron popovi.Archimyrmex wedmannaeDlussky G.M. 2012 New Fossil Ants of the Subfamily Myrmeciinae (Hymenoptera, Formicidae) from Germany Paleontological Journal 46 3 288–292 10.1134/s0031030111050054 2012PalJ...46..288D 83891156 Sp. novValidDlusskyMiddle Eocene Messel PitGermanyA myrmeciine ant.thumb|center|150px|Archimyrmex wedmannaeAnthophorula (Anthophorula) persephoneMichael S. Engel David A. Grimaldi Victor H. González Ismael A. Hinojosa-Díaz Charles D. Michener 2012 An exomalopsine bee in early Miocene amber from the Dominican Republic (Hymenoptera, Apidae) American Museum Novitates 3758 1–16 10.1206/3758.2 2246/6371 82179983 https://www.biodiversitylibrary.org/item/276416 Sp. novValidEngel in Engel et al.Early MioceneDominican RepublicAn exomalopsine apid bee found in Dominican amber, a species of Anthophorula.AtefiaLars Krogmann Michael S. Engel Günter Bechly André Nel 2013 Lower Cretaceous origin of long-distance mate finding behaviour in Hymenoptera (Insecta) Journal of Systematic Palaeontology 11 1 83–89 10.1080/14772019.2012.693954 2013JSPal..11...83K 84070677 Gen. et sp. novValidKrogmann et al.Early CretaceousCrato FormationBrazilA tenthredinoid hymenopteran sharing morphological characters with the families Pergidae and Diprionidae. The type species is Atefia rasnitsyni.BoltonidrisRadchenko Dlussky G.M. 2012 Boltonidris Gen. Nov., the First Extinct Stenammini Ant Genus (Hymenoptera, Formicidae) from the Late Eocene Rovno Amber Annales Zoologici 62 4 627–631 10.3161/000345412X659687 84291016 Gen. et sp. novValidRadchenko & DlusskyLate EoceneRovno amberUkraineA stenammine myrmicine ant found in Rovno amber. The type species is Boltonidris mirabilis.Bombus (Bombus) randeckensisTorsten Wappler Thibaut De Meulemeester A. Murat Aytekin Denis Michez Michael S. Engel 2012 Geometric morphometric analysis of a new Miocene bumble bee from the Randeck Maar of southwestern Germany (Hymenoptera: Apidae) Systematic Entomology 37 4 784–792 10.1111/j.1365-3113.2012.00642.x 2012SysEn..37..784W 84979697 Sp. novValidWappler & Engel in Wappler et al.MioceneGermanyA bumblebee.Casaleia eocenica Dlussky G.M. Wedmann S. The poneromorph ants (Hymenoptera, Formicidae: Amblyoponinae, Ectatomminae, Ponerinae) of Grube Messel, Germany: High biodiversity in the Eocene Journal of Systematic Palaeontology 10 4 725–753 2012 10.1080/14772019.2011.628341 2012JSPal..10..725D 83928415 Taylor & FrancisDlussky G.M. Putyatina T.S. 2014 Early Miocene ants (Hymenoptera, Formicidae) from Radoboj, Croatia Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen 272 3 237–285 10.1127/0077-7749/2014/0409 2014NJGPA.272..237D Sp novvalidDlussky & WedmannLutetian Messel FormationGermanyAn amblyoponine ant.thumb|center|150px|Casaleia eocenicaCascoscelioPoinar Jr. George Buckley Ron 2011 Predatory behaviour of the social orb-weaver spider, Geratonephila burmanica n. gen., n. sp. (Araneae: Nephilidae) with its wasp prey, Cascoscelio incassus n. gen., n. sp. (Hymenoptera: Platygastridae) in Early Cretaceous Burmese amber Historical Biology: An International Journal of Paleobiology 24 5 519–525 10.1080/08912963.2011.640399 2012HBio...24..519P 86838688 Gen. et sp. nov. ValidPoinar in Poinar & BuckleyEarly CretaceousBurmese amberMyanmarA platygastrid wasp, type species is C. incassus.thumb|center|upright|Cascoscelio incassaCephaloponeGen. et 2 sp. novValidDlussky & WedmannEoceneMessel pitGermanyA poneromorph ant. Two species: C. potens & C. grandis.thumb|center|150px|Cephalopone grandisCratoenigmaLars Krogmann Andre Nél 2012 On the edge of parasitoidism: a new Lower Cretaceous woodwasp forming the putative sister group of Xiphydriidae + Euhymenoptera Systematic Entomology 37 1 215–222 10.1111/j.1365-3113.2011.00608.x 2012SysEn..37..215K 85251731 Gen. et sp. novValidKrogmann & NélEarly Cretaceous (Aptian)Crato FormationBrazilA probable relative of xiphydriids and euhymenopterans. The type species is C. articulata.Cretevania soplaensisRicardo Pérez-De La Fuente Enrique Peñalver Jaime Ortega-Blanco 2012 A new species of the diverse Cretaceous genus Cretevania Rasnitsyn, 1975 (Hymenoptera: Evaniidae) from Spanish amber Zootaxa 3514 70–78 https://www.mapress.com/zootaxa/2012/f/z03514p078f.pdf 10.11646/zootaxa.3514.1.4 10261/277032 Sp. novValidDe La Fuente, Peñalver & Ortega-BlancoAlbianSpainAn evaniid, a species of Cretevania.CretorhyssalusGen. et sp. novValidBelokobylskijCenomanian Ola FormationRussiaA braconid wasp. Type species is Cretorhyssalus brevis.CyrtoponeGen. et 4 sp. novValidDlussky & WedmannEoceneMessel FormationGermanyA ponerinae ant. Genus contains four species: Cyrtopone curiosa, C. elongata, C. microcephala and C. striata.thumb|center|150px|C. striataGulgongaStefanie K. Oberprieler Alexandr P. Rasnitsyn Denis J. Brothers 2012 The first wasps from the Upper Jurassic of Australia (Hymenoptera: Evanioidea, Praeaulacidae) Zootaxa 3503 47–54 https://www.mapress.com/zootaxa/2012/f/z03503p054f.pdf 10.11646/zootaxa.3503.1.3 Gen. et sp. novValidOberprieler, Rasnitsyn & BrothersLate JurassicTalbragar Fish BedAustraliaA praeaulacid evanioid. The type species is Gulgonga beattiei.Haidomyrmex scimitarusBarden P. Grimaldi D. 2012 Rediscovery of the bizarre Cretaceous ant Haidomyrmex Dlussky (Hymenoptera: Formicidae), with two new species American Museum Novitates 3755 1–16 10.1206/3755.2 http://digitallibrary.amnh.org/dspace/bitstream/handle/2246/6368/N3755.pdf?sequence=1 2246/6368 83598305 2013-10-22 2013-04-23 https://web.archive.org/web/20130423110550/http://digitallibrary.amnh.org/dspace/bitstream/handle/2246/6368/N3755.pdf?sequence=1 dead Sp. novValidBarden & GrimaldiLate Aptian to Early CenomanianBurmese amberMyanmarA Sphecomyrmin antthumb|center|150px|Haidomyrmex scimitarusHaidomyrmex zigrasiSp. novValidBarden & GrimaldiCampanianBurmese amberMyanmarA Sphecomyrmin antthumb|center|150px|Haidomyrmex zigrasiHyptia deansiJohn T. Jennings Lars Krogmann Steven L. Mew 2012 Hyptia deansi sp. nov., the first record of Evaniidae (Hymenoptera) from Mexican amber Zootaxa 3349 63–68 https://www.mapress.com/zootaxa/2012/f/zt03349p068.pdf 10.11646/zootaxa.3349.1.7 Sp. novValidJennings, Krogmann & MewLate Oligocene or early Miocene MexicoAn evaniid hymenopteran, a species of Hyptia.Ibalia (Ibalia) electra Michael S. Engel Zhiwei Liu 2012 A New Species of Ibaliid Wasp in Baltic Amber (Hymenoptera: Ibaliidae) Journal of the Kansas Entomological Society 85 2 164–168 10.2317/JKES120421.1 85237154 Sp. novValidEngel & LiuEoceneEuropeAn ibaliid found in Baltic amber, a species of Ibalia.LancepyrisCelso O. Azevedo Dany Azar 2012 A new fossil subfamily of Bethylidae (Hymenoptera) from the Early Cretaceous Lebanese amber and its phylogenetic position Zoologia (Curitiba) 29 3 210–218 10.1590/S1984-46702012000300004 free Gen. et sp. novValidAzevedo & AzarEarly CretaceousLebanonA lancepyrine bethylid found in Early Cretaceous Lebanese amber. The type species is Lancepyris opertus.Lonchodryinus balticusMassimo Olmi Adalgisa Guglielmino 2012 A contribution to the knowledge of Dryinidae from Late Eocene Baltic amber (Hymenoptera: Chrysidoidea), with description of new subfamily, Protodryininae subfam. nov. Zootaxa 3351 15–26 https://www.mapress.com/zootaxa/2012/f/z03351p026f.pdf 10.11646/zootaxa.3351.1.2 Sp. novValidOlmi & GuglielminoLate EoceneEuropeA dryinid found in Baltic amber, a species of Lonchodryinus.MagadanobraconSergey A. Belokobylskij 2012 Cretaceous braconid wasps from the Magadan Province of Russia Acta Palaeontologica Polonica 57 2 351–361 https://app.pan.pl/article/item/app20100120.html 10.4202/app.2010.0120 free Gen. et 2 sp. novValidBelokobylskijCenomanian Ola FormationRussiaA braconid wasp. The type species is M. rasnitsyni; a second species is M. zherikhini.MesseleponeGen. et sp. novValidDlussky & WedmannEoceneMessel pitGermanyA poneromorph ant. The type species is M. leptogenoides.thumb|center|150px|Messelepone leptogenoidesMirabythusYaping Cai Yunyun Zhao Chungkun Shih Dong Ren 2012 A new genus of Scolebythidae (Hymenoptera: Chrysidoidea) from the Early Cretaceous of China Zootaxa 3504 56–66 https://www.mapress.com/zootaxa/2012/f/z03504p066f.pdf 10.11646/zootaxa.3504.1.2 Gen. et 2 sp. novValidCai, Shih & Ren in Cai et al.Early CretaceousYixian FormationChinaA scolebythid chrysidoid. The genus contains two species: Mirabythus lechrius and M. liae.Pachycondyla eocenicaSp. novValidDlussky & WedmannEoceneMessel pitGermanyA Ponerinae ant.thumb|center|150px| Pachycondyla eocenicaPachycondyla lutziSp. novValidDlussky & WedmannEoceneMessel pitGermanyA ponerine antthumb|center|150px|Pachycondyla lutziPachycondyla? messelianaSp. novValidDlussky & WedmannEoceneMessel pitGermanyA ponerine ant.thumb|center|150px|Pachycondyla? messelianaPachycondyla minutaDlussky G. M. Rasnitsyn A. P. Perfilieva K. S. 2015 The ants (Hymenoptera: Formicidae) of Bol'shaya Svetlovodnaya (late Eocene of Sikhote-Alin, Russian far east) Caucasian Entomological Bulletin 11 1 131–152 10.23885/1814-3326-2015-11-1-131-152 free Sp. novsynonymDlussky & WedmannEoceneMessel pitGermanyA ponerine ant, name occupied, renamed P. parvula.thumb|center|150px|Pachycondyla parvulaPachycondyla petiolosaSp. novValidDlussky & WedmannEoceneMessel pitGermanyA ponerine ant.thumb|center|150px|Pachycondyla petiolosaPachycondyla petrosaSp. novValidDlussky & WedmannEoceneMessel pitGermanyA poneromorph ant, a species of Pachycondyla.thumb|center|150px|Pachycondyla petrosaPlatyxyelaMei Wang Chungkun Shih Dong Ren 2012 Platyxyela gen. nov. (Hymenoptera, Xyelidae, Macroxyelinae) from the Middle Jurassic of China Zootaxa 3456 82–88 https://www.mapress.com/zootaxa/2012/f/z03456p088f.pdf 10.11646/zootaxa.3456.1.4 Gen. et sp. novValidWang, Shih & RenMiddle JurassicJiulongshan FormationChinaA macroxyeline xyelid sawfly. The type species is Platyxyela unica.Praeichneumon dzhidensisD. S. Kopylov 2012 New species of Praeichneumonidae (Hymenoptera, Ichneumonoidea) from the Lower Cretaceous of Transbaikalia Paleontological Journal 46 1 66–72 10.1134/S0031030112010078 2012PalJ...46...66K 84409422 sp novValidKopylovLower Cretaceous RussiaA praeichneumonid ichneumonoid, a species of Praeichneumon.Praeichneumon khamardabanicussp novValidKopylovLower Cretaceous RussiaA praeichneumonid ichneumonoid, a species of Praeichneumon.Praeichneumon zakhaaminicussp novValidKopylovLower Cretaceous RussiaA praeichneumonid ichneumonoid, a species of Praeichneumon.Prionomyrmex wappleriSp. novValidDlusskyLate Oligocene GermanyA myrmeciine ant.thumb|center|150px|Prionomyrmex wappleriProtodryinusGen. et sp. novValidOlmi & GuglielminoLate EoceneEuropeA dryinid found in Baltic amber. The type species is Protodryinus eocenicus.Protopone? dubiaSp. novValidDlussky & WedmannEoceneMessel FormationGermanyA ponerine antthumb|center|150px|Protopone? dubiaProtopone germanicaSp. novValidDlussky & WedmannEoceneMessel pitGermanyA ponerine antthumb|center|150px|Protopone germanicaProtopone magnaSp. novValidDlussky & WedmannEoceneMessel pitGermanyA ponerine antthumb|center|150px|Protopone magnaProtopone oculataSp. novValidDlussky & WedmannEoceneMessel pitGermanyA ponerine antthumb|center|150px|Protopone oculataProtopone sepultaSp. novValidDlussky & WedmannEoceneMessel pitGermanyA ponerine antthumb|center|150px|Protopone sepultaProtopone vetulaSp. novValidDlussky & WedmannEoceneMessel pitGermanyA ponerine antthumb|center|150px|Protopone vetulaPseudectatommaGen. et 2 sp. novValidDlussky & WedmannEoceneMessel pitGermanyAn Ectatommin ant. Two species: P. eocenica and P. striatula.thumb|center|150px|Pseudectatomma eocenicaTrichelyonJaime Ortega-Blanco Hukam Singh Michael S. Engel 2012 First amber fossil Rhysipolini (Hymenoptera: Braconidae): a new genus and species in Early Eocene Cambay amber Acta Entomologica Musei Nationalis Pragae 52 2 585–594 https://www.aemnp.eu/data/article-1391/1372-52_2_585.pdf Gen. et sp. novDisputedOrtega-Blanco & Engel in Ortega-Blanco, Singh & EngelEarly Eocene (Ypresian)IndiaA rhysipoline exothecine braconid wasp found in Cambay amber. The type species is Trichelyon tadkeshwarense. Belokobylskij & Manukyan (2024) considered Trichelyon to be a junior synonym of the genus Clinocentrus, though the authors maintained T. tadkeshwarense as a distinct species within the latter genus, resulting in a new combination Clinocentrus tadkeshwarense.Sergey A. Belokobylskij Andranik R. Manukyan 2024 First reliable fossil record of the subfamily Rhysipolinae (Hymenoptera: Braconidae): a new subgenus and species of the genus Rhysipolis Foerster, 1863 from Baltic amber Zootaxa 5448 4 591–600 10.11646/zootaxa.5448.4.10 39646235 Urotryphon baikurensisSp. novValidKopylovLate CretaceousRussiaAn ichneumonid, a species of Urotryphon.Zatania electraJohn S. Lapolla Robert J. Kallal Seán G. Brady 2012 A new ant genus from the Greater Antilles and Central America, Zatania (Hymenoptera: Formicidae), exemplifies the utility of male and molecular character systems Systematic Entomology 37 1 200–214 10.1111/j.1365-3113.2011.00605.x 2012SysEn..37..200L 84919642 Sp. novValidLapolla, Kallal & BradyEocene or MioceneDominican amberDominican RepublicA formicine ant, a species of Zatania.thumb|center|upright|Zatania electra Mecoptera Name Novelty Status Authors Age Unit Location Notes ImagesBaltipanorpaKrzemiński W. Soszyńska-Maj A. 2012 A new genus and species of scorpionfly (Mecoptera) from Baltic amber, with an unusually developed postnotal organ Systematic Entomology 37 1 223–228 10.1111/j.1365-3113.2011.00602.x 2012SysEn..37..223K 83472551 Gen. et sp. novValidKrzemiński & Soszyńska-MajMiddle EoceneBaltic amberPolandA panorpid scorpionfly. The type species is B. damzeni.ExilibittacusXiaoguang Yang Dong Ren Chungkun Shih 2012 New fossil hangingflies (Mecoptera, Raptipeda, Bittacidae) from the Middle Jurassic to Early Cretaceous of Northeastern China Geodiversitas 34 4 785–799 10.5252/g2012n4a4 2012Geodv..34..785Y 44179968 http://sciencepress.mnhn.fr/en/periodiques/geodiversitas/34/4/de-nouveaux-fossiles-de-mouches-scorpion-mecoptera-raptipeda-bittacidae-du-jurassique-moyen-au-cretace-inferieur-du-nord-est-de-la-chine subscription Gen. et sp. novValidYang, Ren & ShihMiddle JurassicJiulongshan FormationChinaA bittacid hangingfly. The type species is E. lii.thumb|center|upright|Exilibittacus liiJeholopsyche bellaXiao Qiao Chungkun Shih Dong Ren 2012 Three new species of aneuretopsychids (Insecta: Mecoptera) from the Jehol Biota, China Cretaceous Research 36 146–150 10.1016/j.cretres.2012.03.004 2012CrRes..36..146Q Sp. novValidQiao, Shih & RenEarly CretaceousYixian FormationChinaAn aneuretopsychid mecopteran.Jeholopsyche completaSp. novValidQiao, Shih & RenEarly CretaceousYixian FormationChinaAn aneuretopsychid mecopteran.Jeholopsyche maximaSp. novValidQiao, Shih & RenEarly CretaceousYixian FormationChinaAn aneuretopsychid mecopteran.JuracimbrophlebiaYongjie Wang Conrad C. Labandeira Chungkun Shih Qiaoling Ding Chen Wang Yunyun Zhao Dong Ren 2012 Jurassic mimicry between a hangingfly and a ginkgo from China Proceedings of the National Academy of Sciences of the United States of America 109 50 20514–20519 10.1073/pnas.1205517109 23184994 3528590 2012PNAS..10920514W free Gen. et sp. novValidWang, Labandeira, Shih & Ren in Wang et al.Middle Jurassic (Bathonian or Callovian)Jiulongshan FormationChinaA cimbrophlebiid mecopteran. Adults with Yimaia ginkglaean leaf mimicry. The type species is J. ginkgofolia.thumb|center|upright|Juracimbrophlebia ginkgofoliaMegabittacus spatiosusSp. novValidYang, Ren & ShihLate Jurassic or Early CretaceousYixian FormationChinaA hangingfly, a species of Megabittacus.Orthophlebia nervulosaXiao Qiao Chungkun Shih Dong Ren 2012 Two new Middle Jurassic species of orthophlebiids (Insecta: Mecoptera) from Inner Mongolia, China Alcheringa: An Australasian Journal of Palaeontology 36 4 467–472 10.1080/03115518.2012.671689 2012Alch...36..469Q 129380801 sp novValidQiao, Shih & RenMiddle Jurassic Jiulongshan FormationChinaA mecopteran, a species of Orthophlebia.Orthophlebia riccardiiJulián F. Petrulevičius Dong Ren 2012 A new species of "Orthophlebiidae" (Insecta: Mecoptera) from the Middle Jurassic of Inner Mongolia, China Revue de Paléobiologie, Genève spécial 11 311–315 http://www.ville-ge.ch/mhng/paleo/paleo-pdf/spec-11/pal-spec-11-21.pdf Sp. novValidPetrulevičius & RenMiddle JurassicJiulongshan FormationChinaA mecopteran. Originally described as a species of Orthophlebia, subsequently transferred to the separate genus Gigaphlebia.Agnieszka Soszyńska-Maj Wiesław Krzemiński Katarzyna Kopeć Yizi Cao Dong Ren 2018 Large Jurassic scorpionflies belonging to a new subfamily of the family Orthophlebiidae (Mecoptera) Annales Zoologici 68 1 85–92 10.3161/00034541ANZ2018.68.1.004 90298406 Orthophlebia stigmosasp novValidQiao, Shih & RenMiddle Jurassic Jiulongshan FormationChinaA mecopteran. Originally described as a species of Orthophlebia, subsequently transferred to the separate genus Longiphlebia.PerfecticimbrophlebiaXiao-Guang Yang Chung-Kun Shih Dong Ren Julián F Petrulevičius 2012 New Middle Jurassic hangingflies (Insecta: Mecoptera) from Inner Mongolia, China Alcheringa: An Australasian Journal of Palaeontology 36 2 195–201 10.1080/03115518.2012.622143 2012Alch...36..195Y 128687609 Gen. et sp. novValidYang, Shih & Ren in Yang et al.Middle Jurassic Jiulongshan FormationChinaA cimbrophlebiid mecopteran. The type species is Perfecticimbrophlebia laetus.Preanabittacus validussp novValidYang, Shih & Ren in Yang et al.Middle Jurassic Jiulongshan FormationChinaA hangingfly, a species of Preanabittacus. Neuroptera Name Novelty Status Authors Age Unit Location Notes ImagesAlloepipsychopsisVladimir N. Makarkin Qiang Yang YuanYuan Peng Dong Ren 2012 A comparative overview of the neuropteran assemblage of the Lower Cretaceous Yixian Formation (China), with description of a new genus of Psychopsidae (Insecta: Neuroptera) Cretaceous Research 35 57–68 10.1016/j.cretres.2011.11.013 2012CrRes..35...57M Gen. et sp. novValidMakarkin, Yang, et al.BarremianYixian FormationChinaA psychopsid. Type species is Alloepipsychopsis lata.ChorilingiaChao-Fan Shi Yong-Jie Wang Qiang Yang Dong Ren 2012 Chorilingia (Neuroptera: Grammolingiidae): a new genus of lacewings with four species from the Middle Jurassic of Inner Mongolia, China Alcheringa: An Australasian Journal of Palaeontology 36 3 309–318 10.1080/03115518.2012.644994 2012Alch...36..309S 84255854 Gen. et 4 sp. novValidShi et al.Middle JurassicJiulongshan FormationChinaA grammolingiid neuropteran. Genus contains four species: of C. euryptera, C. parvica, C. translucida and C. peregrina.GuyilingChaofan Shi Olivier Béthoux Chungkun Shih Dong Ren 2012 Guyiling jianboni gen. et sp.n., an antlion-like lacewing, illuminating homologies and transformations in Neuroptera wing venation Systematic Entomology 37 3 617–631 10.1111/j.1365-3113.2012.00633.x 2012SysEn..37..617S 84467796 Gen. et sp. novValidShi et al.Early CretaceousYixian FormationChinaA member of Myrmeleontiformia of uncertain phylogenetic placement. The type species is G. jianboni.HallucinochrysaRicardo Pérez-de la Fuente Xavier Delclòs Enrique Peñalver Mariela Speranza Jacek Wierzchos Carmen Ascaso Michael S. Engel 2012 Early evolution and ecology of camouflage in insects Proceedings of the National Academy of Sciences of the United States of America 109 52 21414–21419 10.1073/pnas.1213775110 23236135 3535654 2012PNAS..10921414P free Gen. et sp. novValidPérez-de la Fuente, Delclòs, Peñalver & Engel in Pérez-de la Fuente et al.Early Cretaceous (Albian)SpainA chrysopoid neuropteran (a relative of the green lacewings). The type species is Hallucinochrysa diogenesi.KareninoidesQiang Yang Vladimir N. Makarkin Dong Ren 2012 New fossil Mesochrysopidae (Neuroptera) from the Mesozoic of China Zootaxa 3597 1–14 https://www.mapress.com/zootaxa/2012/f/z03597p014f.pdf 10.11646/zootaxa.3597.1.1 Gen. et sp. et comb. novValidYang, Makarkin & RenEarly CretaceousLa Huerguina Formation Yixian FormationChina SpainA mesochrysopid neuropteran. The type species is Kareninoides lii; genus might also contain "Armandochrysopa" inexpecta Nel et al. (2005).Leptolingia oblongaAlexander V. Khramov 2012 The new fossil lacewings of Grammolingiidae (Neuroptera) from the Jurassic of Central Asia and Mongolia, with notes on biogeography of the family Zootaxa 3478 297–308 10.11646/zootaxa.3478.1.28 https://www.mapress.com/zootaxa/2012/f/z03478p308f.pdf Sp. novValidKhramovLate JurassicMongoliaA grammolingiid neuropteran, a species of Leptolingia.Litholingia longaSp. novValidKhramovEarly or Middle JurassicKyrgyzstanA grammolingiid neuropteran, a species of Litholingia.ParakseneuraQiang Yang Vladimir N. Makarkin Shaun L. Winterton Alexander V. Khramov Dong Ren 2012 A Remarkable New Family of Jurassic Insects (Neuroptera) with Primitive Wing Venation and Its Phylogenetic Position in Neuropterida PLOS ONE 7 9 e44762 10.1371/journal.pone.0044762 23028608 3445537 2012PLoSO...744762Y free Gen. et 11 sp. novValidYang, Makarkin & Ren in Yang et al.Middle Jurassic Jiulongshan FormationChinaA parakseneurid neuropteran. Genus contains 11 species.Protoaristenymphes daohugouensisSp. novValidYang, Makarkin & RenMiddle JurassicDaohugou BedsChinaA mesochrysopid neuropteran, a species of Protoaristenymphes.ProtolingiaGen. et sp. novValidKhramovEarly or Middle JurassicKyrgyzstanA grammolingiid neuropteran. The type species is Protolingia mira.PseudorapismaGen. et 3 sp. novValidYang, Makarkin & Ren in Yang et al.Middle Jurassic Jiulongshan FormationChinaA parakseneurid neuropteran. Genus contains 3 species: P. jurassicum, P. maculatum and P. angustipenne.ShuraboneuraGen. et sp. novValidKhramov & Makarkin in Yang et al.Early or Middle Jurassic Sogul FormationKyrgyzstanA parakseneurid neuropteran. The type species is Shuraboneura ovata. Notoptera Name Novelty Status Authors Age Unit Location Notes ImagesDuoduoYingying Cui 2012 New data on the Blattogryllidae-Plesioblattogryllidae-Grylloblattidae complex (Insecta: Grylloblattida: Blattogryllopterida tax.n.) Arthropod Systematics & Phylogeny 70 3 167–180 10.3897/asp.70.e31760 free Gen. et sp. novJunior synonymCuiMiddle JurassicDaohugou BedsChinaA putative relative of grylloblattids. The type species is Duoduo qianae. Huang & Nel (2023) considered the genus Duoduo to be a junior synonym of the genus Plesioblattogryllus,Di-Ying Huang André Nel A third species of Plesioblattogryllus Huang et al., 2008 discovered in the Middle-Upper Jurassic Daohugou Beds (Insecta: Plesioblattogryllidae) 2023 Palaeoentomology 6 2 191–197 10.11646/palaeoentomology.6.2.10 2023Plegy...6.2.10H 258405260 while Cui et al. (2023) considered the species D. qianae to be a junior synonym of the species Plesioblattogryllus magnificus.Yingying Cui Olivier Béthoux Nan Yang Dong Ren New data and systematic considerations on the Blattogryllopterida (Insecta: Grylloblattida) from the Jurassic Daohugou locality in China 2023 Palaeoentomology 6 2 124–132 10.11646/palaeoentomology.6.2.4 2023Plegy...6..2.4C 258399002 KitshugaD. S. Aristov 2012 Composition and distribution of the family Cacurgidae (Insecta; Grylloblattida) Paleontological Journal 46 3 250–257 10.1134/S0031030112030033 2012PalJ...46..250A 85111191 Gen. et sp. novValidAristovUpper Permian RussiaA cacurgid grylloblattid. The type species is Kitshuga ryzhkovae.Oborella brauckmanniJakub Prokop Klaus-Dieter Weiß Roger-Paul Dechambre Roger-Paul Dechambre André Nel 2012 Early Permian insects from Saar-Nahe Basin of Odernheim town site, Rheinland-Pfalz in Germany (Insecta, Grylloblattida, Blattinopseida) Geodiversitas 34 2 271–281 http://sciencepress.mnhn.fr/en/periodiques/geodiversitas/34/2/nouveaux-insectes-fossiles-insecta-grylloblattida-blattinopseida-du-permien-inferieur-du-bassin-de-saar-nahe-odernheim-ville-allemagne 10.5252/g2012n2a2 2012Geodv..34..271P 130094576 subscription sp novValidProkop et al.Early Permian Meisenheim FormationGermanyA euryptilonid grylloblattid, a species of Oborella.Oborella germanicasp novValidProkop et al.Early Permian Meisenheim FormationGermanyA grylloblattid. Originally described as a species of Oborella, but subsequently transferred to the genus Pictoborella.Yingying Cui Stephan Brauner Joerg W. Schneider Olivier Béthoux 2021 Grylloblattidan insects from Sperbersbach and Cabarz (Germany), two new early Permian and insect-rich localities Journal of Paleontology 96 2 355–374 10.1017/jpa.2021.101 243486826 free Odonatoptera Name Novelty Status Authors Age Unit Location Notes ImagesAustroprotolindeniaBeattie R.G. Nel A. 2012 A new dragonfly, Austroprotolindenia jurassica (Odonata: Anisoptera), from the Upper Jurassic of Australia Alcheringa: An Australasian Journal of Palaeontology 36 2 189–193 10.1080/03115518.2011.598055 2012Alch...36..189B 83651613 Gen. et sp. novValidBeattie & NelLate Jurassic Talbragar Fossil Fish BedAustraliaA dragonfly. The type species is Austroprotolindenia jurassica.AzaroneuraAndré Nel Günter Bechly Jakub Prokop Olivier Béthoux Gunther Fleck 2012 Systematics and evolution of Paleozoic and Mesozoic damselfly-like Odonatoptera of the 'protozygopteran' grade Journal of Paleontology 86 1 81–104 10.1666/11-020.1 Gen. et sp. novValidNel et al.Permian (Kungurian)Koshelevka FormationRussiaA protozygopteran. The type species is A. permiana.BalticoagrionGünter Bechly 2012 An interesting new fossil relict damselfly (Odonata: Zygoptera: Coenagrionoidea) from Eocene Baltic amber Palaeodiversity 5 51–55 https://www.palaeodiversity.org/pdf/05/Palaeodiversity_5_51-55_Bechly.pdfGen. et sp. novValidBechlyEoceneEuropeA damselfly found in Baltic amber. The type species is Balticoagrion paulyi.DecoraeshnaYongjun Li André Nel Dong Ren Hong Pang 2012 New gomphaeschnids and progobiaeshnids from the Yixian Formation in Liaoning Province (China) illustrate the tremendous Upper Mesozoic diversity of the aeshnopteran dragonflies Geobios 45 4 339–350 10.1016/j.geobios.2011.11.016 2012Geobi..45..339L Gen. et sp. novValidLi et al.Early Cretaceous Yixian FormationChinaA progobiaeshnid dragonfly. The type species is Decoraeshna preciosus.EngellestesGen. et sp. novValidNel et al.Permian (Kungurian)Koshelevka FormationRussiaA protozygopteran. The type species is E. chekardensis.Epilestes angustapterixSp. novValidNel et al.Late PermianRussiaA protozygopteran.GermanostenolestesAndré Nel Gunther Fleck 2012 A new genus and species of Sieblosiidae from the Middle Miocene of Germany (Odonata: Epiproctophora) Zootaxa 3582 64–68 https://www.mapress.com/zootaxa/2012/f/z03582p068f.pdf 10.11646/zootaxa.3582.1.7 Gen. et sp. novValidNel & FleckMiddle MioceneGermanyA sieblosiid epiproctophoran odonate. The type species is Germanostenolestes lutzi.Karatawia sinicaYong-Jun Li André Nel Dong Ren Bing-Lan Zhang Hong Pang 2012 Reassessment of the Jurassic damsel-dragonfly genus Karatawia (Odonata: Campterophlebiidae) Zootaxa 3417 64–68 https://www.mapress.com/zootaxa/2012/f/z03417p068f.pdf 10.11646/zootaxa.3417.1.4 Sp. novValidLi, Nel & Ren in Li et al.Middle JurassicJiulongshan FormationChinaA campterophlebiid odonate,a species of Karatawia.Kennedya azariSp. novValidNel et al.Permian (Kungurian)Koshelevka FormationRussiaA protozygopteran.Kennedya ivensisSp. novValidNel et al.Late PermianRussiaA protozygopteran.Kennedya pritykinaeSp. novValidNel et al.Late PermianRussiaA protozygopteran.LuiseiaGen. et sp. novValidNel et al.Carboniferous/Permian boundaryBursum FormationUnited StatesA protozygopteran. The type species is L. breviata.Mongoliaeshna exiguusensSp. novValidLi et al.Early Cretaceous Yixian FormationChinaA progobiaeshnid dragonfly, a species of Mongoliaeshna.Mongoliaeshna hadrensSp. novValidLi et al.Early Cretaceous Yixian FormationChinaA progobiaeshnid dragonfly, a species of Mongoliaeshna.OligomazonGen. et comb. novValidZessin & BrauckmannCarboniferous (Pennsylvanian)United StatesA member of Meganisoptera, a paralogid; a new genus for "Oligotypus" makowskii Carpenter & Richardson (1971).ParagilsoniaYa Su Zhijun Zhang Youchong Hong 2013 Two new ancient griffenflies (Insecta: Odonatoptera) from the Namurian of Ningxia, China Insect Systematics & Evolution 43 1 1–10 10.1163/187631212X624205 Gen. et sp. nov (but see notes)ValidZhang, Hong & Su in Su et al.Carboniferous (Namurian)ChinaA member of Odonatoptera; the type species is Paragilsonia orientalis. Li et al. (2013) synonymized Paragilsonia with the genus Tupus Sellards (1906); however, they retained P. orientalis as a distinct species within the latter genus.Yongjun Li Olivier Béthoux Hong Pang Dong Ren 2013 Early Pennsylvanian Odonatoptera from the Xiaheyan locality (Ningxia, China): new material, taxa, and perspectives Fossil Record 16 1 117–139 10.1002/mmng.201300006 free 2013FossR..16..117L ParalogoboraWolfgang Zessin Carsten Brauckmann 2012 Eine neue fossile Libelle: Paralogobora guentherpetersi gen. nov., sp. nov. aus dem Unterperm von Obora, Tschechische Republik, und eine neue Gattung: Oligomazon gen. nov. für Oligotypus makowskii (Meganisoptera: Paralogidae) Libellula: Zeitschrift der Gesellschaft Deutschsprachiger Odonatologen (GdO) E.V. Supplement 12 237–245 Gen. et sp. novValidZessin & BrauckmannEarly PermianCzech RepublicA member of Odonata. The type species is Paralogobora guentherpetersi.Permolestes sheimogoraiSp. novValidNel et al.Late PermianRussiaA protozygopteran.Permolestes soyanaiensisSp. novValidNel et al.Late PermianRussiaA protozygopteran.Progoneura grimaldiiSp. novValidNel et al.Early PermianWellington FormationUnited StatesA protozygopteran.Pseudocymatophlebia bodaYongjun Li Gang Han André Nel Dong Ren Hong Pang Xueling Liu 2012 A new fossil petalurid dragonfly (Odonata: Petaluroidea: Aktassiidae) from the Cretaceous of China Alcheringa: An Australasian Journal of Palaeontology 36 3 319–322 10.1080/03115518.2012.644995 2012Alch...36..319L 128394986 Sp. novValidLi, Nel & Ren in Li et al.Early CretaceousChinaAn aktassiid odonate, a species of Pseudocymatophlebia.SinierasipteraGen. et sp. nov (but see notes)ValidZhang, Hong & Su in Su et al.Carboniferous (Namurian)ChinaA member of Odonatoptera; the type species is Sinierasiptera jini. Li et al. (2013) synonymized Sinierasiptera with the genus Erasipterella Brauckmann (1983); however, they retained S. jini as a distinct species within the latter genus.Sinojagoria cancellosaSp. novValidLi et al.Early Cretaceous Yixian FormationChinaA gomphaeschnid dragonfly, a species of Sinojagoria.Sinojagoria magnaSp. novValidLi et al.Early Cretaceous Yixian FormationChinaA gomphaeschnid dragonfly, a species of Sinojagoria.Solikamptilon pectinatusSp. novValidNel et al.Late PermianRussiaA protozygopteran. Orthopterida Name Novelty Status Authors Age Unit Location Notes ImagesArchaboilus musicusJun-Jie G Montealegre-Z F Robert D. Engel M.S. Ge-Xia Q. Dong R. 2012 Wing stridulation in a Jurassic katydid (Insecta, Orthoptera) produced low-pitched musical calls to attract females Proceedings of the National Academy of Sciences of the United States of America 109 10 3868–3873 10.1073/pnas.1118372109 22315416 3309752 free Sp. novValidGu, Engel and Ren in Gu et al.Bathonian to Callovian Jiulongshan FormationChinaA haglid orthopteran.EoerianthusAndrej V. Gorochov Conrad C. Labandeira 2012 Eocene Orthoptera from Green River Formation of Wyoming (USA) Russian Entomological Journal 21 4 357–370 10.15298/rusentj.21.4.02 free Gen. et sp. novValidGorochov in Gorochov & LabandeiraEoceneBridgerianGreen River FormationLake UnitaUnited StatesColoradoA monkey grasshopper. The type species is E. eocaenicus.EogryllusGen. et 2 sp. novValidGorochov in Gorochov & LabandeiraEoceneGreen River FormationUnited StatesA Gryllini tribe grylline true cricket. The type species is E. unicolor Genus also tentatively includes E.? elongatus.EomogoplistesGen. et sp. novValidGorochov in Gorochov & LabandeiraEoceneBridgerianGreen River FormationLake UnitaUnited StatesColoradoA Mogoplistini tribe Mogoplistine scaly cricket. The type species is E. longipennis.EotetrixGen. et sp. novValidGorochov in Gorochov & LabandeiraEoceneBridgerianGreen River FormationLake UnitaUnited StatesColoradoA batrachideine pygmy grasshopper. The type species is E. unicornis.EotrellaGen. et sp. novValidGorochov in Gorochov & LabandeiraEoceneBridgerianGreen River FormationLake UnitaUnited StatesColoradoA phalangopsine spider cricket The type species is E. mira.EozaclaGen. et 2 sp. novValidGorochov in Gorochov & LabandeiraEoceneBridgerianGreen River FormationLake UnitaUnited StatesColoradoA phalangopsine spider cricket The type species is E. arachnomorpha Genus also includes E. problematica.Heterologus duyiwuerOlivier Béthoux Jun-Jie Gu Dong Ren 2012 A new Upper Carboniferous stem-orthopteran (Insecta) from Ningxia (China) Insect Science 19 2 153–158 10.1111/j.1744-7917.2011.01468.x 2012InsSc..19..153B 55016213 Sp. novValidBéthoux, Gu & RenLate Carboniferous (Namurian)Tupo FormationChinaAn archaeorthopteran (stem-orthopteran), a species of Heterologus.HirsutgerarusWolfgang Zessin 2012 Eine weitere neue Geraride, Hirsutgerarus saxonicus n. gen. n. sp. (Insecta: Panorthoptera: Geraridae), aus dem Oberkarbon (Stephanium C) von Plötz bei Halle (Deutschland) Virgo, Mitteilungsblatt des Entomologischen Vereins Mecklenburg 15 1 10–17 https://www.entomologie-mv.de/download/virgo-15/virg%2015105%20Neue%20Ploetzgerariden.pdf Gen. et sp. novValidZessinCarboniferous (Stephanian)GermanyA gerarid panorthopteran. The type species is Hirsutgerarus saxonicus.Liassophyllum caiiJun-Jie Gu Ge-Xia Qiao Dong Ren 2012 The first discovery of Cyrtophyllitinae (Orthoptera, Haglidae) from the Middle Jurassic and its morphological implications Alcheringa: An Australasian Journal of Palaeontology 36 1 27–34 10.1080/03115518.2011.576535 2012Alch...36...27G 86763856 sp novValidGu & Ren in Gu, Qiao, & RenBathonian to Callovian Jiulongshan FormationChinaA hagloid orthopteran, originally described as a haglid, but subsequently transferred to the family Tuphellidae;Jun-Jie Gu Xin Yang Rong Huang Guijun Yang Yanli Yue Dong Ren 2021 New species and material of Hagloidea (Insecta, Ensifera) from the Yanliao biota of China ZooKeys 1033 183–190 10.3897/zookeys.1033.63571 33958925 8084857 free 2021ZooK.1033..183G a species of Liassophyllum.Miamia maimaiOlivier Béthoux Junjie Gu Yanli Yue Dong Ren 2012 Miamia maimai n. sp., a new Pennsylvanian stem-orthopteran insect, and a case study on the application of cladotypic nomenclature Fossil Record 15 2 103–113 10.1002/mmng.201200008 free Sp. novValidBéthoux et al.Late Carboniferous (early Pennsylvanian)ChinaA stem–orthopteran insect, a species of Miamia.Protogryllus (Protogryllus) lakshmiRicardo Pérez-de La Fuente Sam W. Heads Ismael A. Hinojosa-Díaz Michael S. Engel 2012 The first record of Protogryllinae from the Jurassic of India (Orthoptera: Protogryllidae) Journal of the Kansas Entomological Society 85 1 53–58 10.2317/JKES111103.1 83496373 Sp. novValidPérez-de La Fuente et al.Middle JurassicKota FormationIndiaA protogryllid grylloid ensiferan, a species of Protogryllus.PteromogoplistesGen. et sp. novValidGorochov in Gorochov & LabandeiraEoceneBridgerianGreen River FormationLake UnitaUnited StatesColoradoA Mogoplistini tribe Mogoplistine scaly cricket. The type species is P. grandisGenus tentatively also includes Pronemobius smithii (1890).Pterotriamescaptor? americanusSp. novValidGorochov in Gorochov & LabandeiraEoceneBridgerianGreen River FormationLake UnitaUnited StatesColoradoA gryllotalpine mole cricket.SunopteraYou-chong Hong Zhen-yu Li 2012 The discovery of a new family Sunopteridae fam. nov. (Insecta: Protorthoptera) from Shaanxi Province, China Geological Bulletin of China 31 5 647–652 http://dzhtb.cgs.cn/gbc/ch/reader/view_abstract.aspx?file_no=20120501&flag=1 Gen. et sp. nov ValidHong & LiMiddle TriassicTongchuan FormationChinaA sunopterid "protorthopteran". The type species is Sunoptera orientalis.TettoraptorGen. et sp. novValidGorochov in Gorochov & LabandeiraEoceneBridgerianGreen River FormationLake UnitaUnited StatesColoradoA Tettigoniid grasshopper of uncertain affiliations. The type species is T. maculatus. Raphidioptera Name Novelty Status Authors Age Unit Location Notes ImagesAlavaraphidiaPérez-de la Fuente R. Peñalver E. Delclòs X. Engel M.S. 2012 Snakefly diversity in Early Cretaceous amber from Spain (Neuropterida, Raphidioptera) ZooKeys 204 1–40 10.3897/zookeys.204.2740 22787417 3391719 free 2012ZooK..204....1P Gen et sp. novValidPérez-de la Fuente et al.Albian Escucha FormationSpainA Mesoraphidiid snakefly. The sole species is A. imperterrita.thumb|center|150px|Alavaraphidia imperterritaAmarantoraphidiaGen et sp. novValidPérez-de la Fuente et al.Albian Escucha FormationSpainA Mesoraphidiid snakefly. The sole species is A. ventolina.thumb|center|150px|Amarantoraphidia ventolinaBaissoptera? cretaceoelectraSp. novValidPérez-de la Fuente et al.Albian Escucha FormationSpainA baissopterid snakefly.NecroraphidiaGen. et sp.ValidPérez-de la Fuente et al.Albian Las Peñosas FormationSpainA Mesoraphidiid snakefly. The only species is N. arcuata.thumb|center|150px|Necroraphidia arcuataStyporaphidia? hispanicasp. novValidPérez-de la Fuente et al.Albian Escucha FormationSpainA Mesoraphidiid snakefly. The second species in Styporaphidia.thumb|center|150px|Styporaphidia? hispanica Other Name Novelty Status Authors Age Unit Location Notes ImagesAnebosRussell Garwood Andrew Ross Daniel Sotty Dominique Chabard Sylvain Charbonnier Mark Sutton Philip J. Withers 2012 Tomographic Reconstruction of Neopterous Carboniferous Insect Nymphs PLOS ONE 7 9 e45779 10.1371/journal.pone.0045779 23049858 3458060 2012PLoSO...745779G free Gen. et sp. novValidGarwood et al.Late PennsylvanianAssise de Montceau FormationFranceA polyneopteran insect nymph of uncertain affinities. The type species is Anebos phrixos.thumb|center|150px|Anebos phrixosBechalaJan-Michael Ilger Carsten Brauckmann 2012 First report of Namurian insects (Palaeodictyoptera; Megasecoptera; "basal Neoptera") from the Küchenberg near Fröndenberg/Ruhr (Germany) Paläontologische Zeitschrift 86 2 161–174 10.1007/s12542-011-0122-0 2012PalZ...86..161I 130403110 Gen. et sp. nov ValidIlger & BrauckmannLate Carboniferous (Namurian)GermanyA member of Odonatoptera, assigned to the new family Bechalidae. Originally interpreted as a megasecopteran.Nel A. Ilger J. M. Brauckmann C. Prokop J. 2012 Bechala sommeri Ilger & Brauckmann, 2012 enlightens the Namurian griffenfly diversity (Insecta: Odonatoptera: Bechalidae) Insect Systematics & Evolution 43 2 161–169 10.1163/1876312X04302003 Type species is Bechala sommeri.BruayaphisAndré Nel Jakub Prokop Patricia Nel Philippe Grandcolas Di-Ying Huang Patrick Roques Eric Guilbert Ondřej Dostál Jacek Szwedo 2012 Traits and evolution of wing venation pattern in paraneopteran insects Journal of Morphology 273 5 480–506 10.1002/jmor.11036 22162020 24593650 Gen. et sp. novValidNel, Szwedo & Roques in Nel et al.Late Carboniferous (Bashkirian/Westphalian)FranceA member of Paraneoptera of uncertain phylogenetic placement. The type species is Bruayaphis oudardi.CucullistrigaD. S. Aristov A. P. Rasnitsyn 2012 Revision of the family Idelinellidae, with a review of the Permian Eoblattida (Insecta) Paleontological Journal 46 1 49–60 10.1134/S0031030112010030 2012PalJ...46...49A 140189151 Gen. et sp. novValidAristov & RasnitsynKungurian RussiaA member of Grylloblattida/Eoblattida belonging to the family Idelinellidae; a relative of ice crawlers or cockroaches.D. S. Aristov 2015 Classification of the order Eoblattida (Insecta: Blattidea) with description of new taxa Far Eastern Entomologist 301 1–56 http://www.biosoil.ru/Files/FEE/00000478.pdf 2018-08-25 2018-08-25 https://web.archive.org/web/20180825180153/http://www.biosoil.ru/Files/FEE/00000478.pdf dead Type species is Cucullistriga cucullata.Darekia sanguineaJakub Prokop Wiesław Krzemiński Ewa Krzemińska Dariusz Wojciechowski 2012 Paoliida, a putative stem-group of winged insects: Morphology of new taxa from the Upper Carboniferous of Poland Acta Palaeontologica Polonica 57 1 161–173 10.4202/app.2010.0064 free Gen. et sp. novValidProkop et al.Late Carboniferous (Langsettian) Upper Silesian Coal BasinPolandA member of Paoliida belonging to the family Paoliidae.EochauliodesXingyue Liu Yongjie Wang Chungkun Shih Dong Ren Ding Yang 2012 Early Evolution and Historical Biogeography of Fishflies (Megaloptera: Chauliodinae): Implications from a Phylogeny Combining Fossil and Extant Taxa PLOS ONE 7 7 e40345 10.1371/journal.pone.0040345 22792287 3391272 2012PLoSO...740345L free Gen. et sp. novValidLiu et al.Middle Jurassic (Bathonian-Callovian)Jiulongshan FormationChinaA fishfly. The type species is Eochauliodes striolatus.EospilopsyllusVincent Perrichot Jean-Claude Beaucournu Jürgen Velten 2012 First extinct genus of a flea (Siphonaptera: Pulicidae) in Miocene amber from the Dominican Republic Zootaxa 3438 54–61 https://www.mapress.com/zootaxa/2012/f/z03438p061f.pdf 10.11646/zootaxa.3438.1.3 Gen. et sp. novValidBeaucournu & Perrichot in Perrichot, Beaucournu & VeltenEarly MioceneDominican RepublicA pulicid flea found in Dominican amber. The type species is Eospilopsyllus kobberti.Geinitzia aristoviYingying Cui Sergey Yu Storozhenko Dong Ren 2012 New and little-known species of Geinitziidae (Insecta: Grylloblattida) from the Middle Jurassic of China, with notes on taxonomy, habitus and habitat of these insects Alcheringa: An Australasian Journal of Palaeontology 36 2 251–261 10.1080/03115518.2012.628806 2012Alch...36..251C 129794581 Sp. novValidCui, Storozhenko & RenMiddle Jurassic Jiulongshan FormationChinaA geinitziid, a member of Polyneoptera/Gryllones belonging to the group Reculida; a species of Geinitzia.GymnopollisthripsEnrique Peñalver Conrad C. Labandeira Eduardo Barrón Xavier Delclòs Patricia Nel André Nel Paul Tafforeau Carmen Soriano 2012 Thrips pollination of Mesozoic gymnosperms Proceedings of the National Academy of Sciences of the United States of America 109 22 8623–8628 10.1073/pnas.1120499109 22615414 3365147 2012PNAS..109.8623P free Gen. et 2 sp. nov.ValidPeñalver, Nel & Nel in Peñalver et al.Early Cretaceous SpainA melanthripid thrip. Genus contains two species: Gymnopollisthrips minor and G. maior.Lethe montanaJacqueline Y. Miller Lee D. Miller Michael A. Ivie 2012 A new lepidopteran fossil from the Canyon Ferry Reservoir deposit in southwestern Montana (Lepidoptera: Nymphalidae: Satyrinae) Bulletin of the Allyn Museum 167 1–12 https://www.floridamuseum.ufl.edu/wp-content/uploads/sites/56/2022/04/AME167.pdf Sp. nov ValidMiller, Miller & IvieOligoceneCanyon Ferry LagerstätteUnited StatesA buttefly, a species of LetheLiadotaulius daohugouensisWu Hao Huang Diying 2012 A new species of Liadotaulius (Insecta: Trichoptera) from the Middle Jurassic of Daohugou, Inner Mongolia Acta Geologica Sinica 86 2 320–324 http://www.geojournals.cn/dzxben/ch/reader/view_abstract.aspx?file_no=201202005&flag=1 10.1111/j.1755-6724.2012.00662.x 2012AcGlS..86..320W 140538821 subscription Sp. nov ValidWu & HuangMiddle JurassicDaohugou BedsChinaA caddisfly, a species of Liadotaulius.NetoxenaJae-Cheon Sohn Conrad Labandeira Donald Davis Charles Mitter 2012 An annotated catalog of fossil and subfossil Lepidoptera (Insecta: Holometabola) of the world Zootaxa 3286 1–132 https://www.mapress.com/zootaxa/2012/f/zt03286p132.pdf 10.11646/zootaxa.3286.1.1 Nom. novValidSohn in Sohn et al.Early Cretaceous (late Aptian)Crato FormationBrazilA lepidopteran, possibly a member of Eolepidopterigidae; a replacement name for Xena Martins-Neto (1999).PseudopulexTai-ping Gao Chung-kun Shih Xing Xu Shuo Wang Dong Ren 2012 Mid-Mesozoic Flea-like Ectoparasites of Feathered or Haired Vertebrates Current Biology 22 8 732–735 10.1016/j.cub.2012.03.012 22445298 free 2012CBio...22..732G Gen. et 2 sp. novValidGao, Shih, & Ren in Gao et al.Middle Jurassic to Early Cretaceous ChinaA flea-like insect. The genus contains 2 species: P. jurassicus and P. magnus.RasstrigaGen. et sp. novValidAristov & RasnitsynLate Carboniferous (Desmoinesian) United StatesA member of Grylloblattida/Eoblattida probably belonging to the family Idelinellidae; a relative of ice crawlers or cockroaches. Its type species is Rasstriga americana.ScutistrigaGen. et sp. novValidAristov & RasnitsynKungurian RussiaA member of Grylloblattida/Eoblattida belonging to the family Idelinellidae; a relative of ice crawlers or cockroaches. Its type species is Scutistriga scutata.SharasialisPonomarenko A. G. 2012 Supposed alderfly larva (Insecta, Megaloptera) from the Jurassic locality Shar-Teg, southwestern Mongolia Paleontological Journal 46 4 378–382 10.1134/S0031030112040090 2012PalJ...46..378P 83718393 Gen. et sp. novValidPonomarenkoLate JurassicMongoliaAn alderfly. The type species is Sharasialis fusiformis.SinoprotodiplatysAndré Nel Cédric Aria Romain Garrouste Alain Waller 2016 Evolution and palaeosynecology of the Mesozoic earwigs (Insecta: Dermaptera) Cretaceous Research 33 1 189–195 10.1016/j.cretres.2011.10.002 Gen. et sp. novValidNel et al.Early CretaceousYixian FormationChinaAn earwig belonging to the family Protodiplatyidae. The type species is Sinoprotodiplatys zhangi.StrigullaGen. et comb. novValidAristov & RasnitsynKungurian RussiaA member of Grylloblattida/Eoblattida belonging to the family Idelinellidae; a relative of ice crawlers or cockroaches. A new genus for "Euryptilon" cuculiophoris (Aristov, 2002).StrudiellaRomain Garrouste Gaël Clément Patricia Nel Michael S. Engel Philippe Grandcolas Cyrille D'Haese Linda Lagebro Julien Denayer Pierre Gueriau Patrick Lafaite Sébastien Olive Cyrille Prestianni André Nel 3 2012 A complete insect from the Late Devonian period Nature 488 7409 82–85 10.1038/nature11281 22859205 2012Natur.488...82G 205229663 Gen. et sp. nov. ValidGarrouste et al.Late DevonianBelgiumThe first complete Late Devonian insect, probably terrestrial. The type species is Strudiella devonica.Zdenekia silesiensisSp. novValidProkop et al.Late Carboniferous (Langsettian) Upper Silesian Coal BasinPolandA member of Paoliida belonging to the family Paoliidae. General research The oldest member of the tiger moth subfamily Arctiinae is noted, but not described from the Klondike Mountain Formation.Toussaint E. F. Condamine F. L. Kergoat G. J. Capdevielle-Dulac C. Barbut J. Silvain J. F. Le Ru B. P. 2012 Palaeoenvironmental shifts drove the adaptive radiation of a noctuid stemborer tribe (Lepidoptera, Noctuidae, Apameini) in the Miocene PLOS ONE 7 7 e41377 10.1371/journal.pone.0041377 22859979 3409182 2012PLoSO...741377T free References Category:2012 in paleontology Category:Paleoentomology Category:Lists of arthropods Category:2010s in paleontology