2009 in archosaur paleontology

none 2009 archosaur paleontology science paleontology paleobotany arthropod paleontology paleoentomology paleomalacology paleoichthyology reptile paleontology mammal paleontology The year 2009 in archosaur paleontology was eventful. Archosaurs include the only living dinosaur group — birds — and the reptile crocodilians, plus all extinct dinosaurs, extinct crocodilian relatives, and pterosaurs. Archosaur paleontology is the scientific study of those animals, especially as they existed before the Holocene Epoch began about 11,700 years ago. The year 2009 in paleontology included various significant developments regarding archosaurs. This article records new taxa of fossil archosaurs of every kind that have been described during the year 2009, as well as other significant discoveries and events related to paleontology of archosaurs that occurred in the year 2009. Crurotarsans Delfino M. Smith T. 2009 A reassessment of the morphology and taxonomic status of 'Crocodylus' depressifrons Blainville, 1855 (Crocodylia, Crocodyloidea) based on the Early Eocene remains from Belgium Zoological Journal of the Linnean Society 156 1 140–167 10.1111/j.1096-3642.2008.00478.x free Newly named crurotarsans Name Status Authors Age Unit Location Notes Images ArmadillosuchusMarinho, T.S. Carvalho, I.S. 2009 An armadillo-like sphagesaurid crocodyliform from the Late Cretaceous of Brazil Journal of South American Earth Sciences 27 1 36–41 10.1016/j.jsames.2008.11.005 2009JSAES..27...36M Valid Marinho CarvalhoLate CretaceousAdamantina FormationBrazilNotosuchian with heavy, armadillo-like body armor consisting of flexible bands and rigid shieldsthumb|150px|center|Armadillosuchus BarcinosuchusJuan Martín Leardi Diego Pol 2009 The first crocodyliform from the Chubut Group (Chubut Province, Argentina) and its phylogenetic position within basal Mesoeucrocodylia Cretaceous Research 30 6 1376–1386 10.1016/j.cretres.2009.08.002 2009CrRes..30.1376L 11336/90736 free Valid Leardi PolAptian – AlbianCerro Barcino FormationArgentinaA peirosaurid CollilongusMagdalena Borsuk−Białynicka Andriej G. Sennikov 2009 Archosauriform postcranial remains from the Early Triassic karst deposits of southern Poland Paleontologica Polonica 65 283–328 http://palaeontologia.pan.pl/PP65/PP65_283-328.pdf 13 August 2010 Valid Borsuk−Białynicka SennikovEarly OlenekianCzatkowice 1PolandPossible rauisuchian Coringasuchus Kellner, A.W.A. Pinheiro, A.E.P. Azevedo, S.A.K. Henriques, D.D.R. de Carvalho, L.B. Oliveira, G.R. 2009 A new crocodyliform from the Alcântara Formation (Cenomanian), Cajual Island, Brazil Zootaxa 2030 49–58 10.11646/zootaxa.2030.1.4 Valid Kellner Pinheiro et al.Kellner, Pinheiro, Azevedo, Henriques, de Carvalho, Oliveira.Early CenomanianAlcântara FormationBrazil DuerosuchusSantiago L.A. and Andrés, L.A. 2009 Nuevo género de Crocodylia del Eoceno medio de la Península Ibérica (Zamora, España): Duerosuchus piscator nov. gen., nov. sp Studia Geologica Salmanticensia 45 2 149–173Valid Santiago AndrésMiddle EoceneSpain HypselorhachisButler, R.J. Barrett, P.M. Abel, R.L. Gower, D.J. 2009 A Possible Ctenosauriscid Archosaur from the Middle Triassic Manda Beds of Tanzania Journal of Vertebrate Paleontology 29 4 1022–1031 10.1671/039.029.0404 2009JVPal..29.1022B 86267617 Valid Butler Barrett et al.Butler, Barrett, Abel, Gower.Middle TriassicManda BedsTanzaniaPossible ctenosauriscid KaprosuchusValid Sereno LarsonUpper CretaceousEchkar FormationNigerUnusual large mahajangasuchid with hypertrophied caniniform teeth and posteriorly projecting hornsthumb|150px|center|Kaprosuchus KemkemiaAndrea Cau Simone Maganuco A new theropod dinosaur, represented by a single unusual caudal vertebra from the Kem Kem Beds (Cretaceous) of Morocco Atti Soc. It. Sci. Nat. Museo Civ. Stor. Nat. Milano 150 II 239–257 2009Valid Cau & MaganucoCenomanianKem Kem BedsMoroccoInitially identified as a theropod dinosaur, but subsequently discovered to be a crocodyliform.Lio, G., Agnolin, F., Cau, A. and Maganuco, S. (2012). "Crocodyliform affinities for Kemkemia auditorei Cau and Maganuco, 2009, from the Late Cretaceous of Morocco." Atti della Società Italiana di Scienze Naturali e del Museo di Storia Naturale di Milano, 153 (I), s. 119–126.KhoratosuchusLauprasert, K.; Cuny, G.; Thirakhupt, K. and Suteethorn, V. 2009. Khoratosuchus jintasakuli gen. et sp. nov., an advanced neosuchian crocodyliform from the Early Cretaceous (Aptian-Albian) of NE Thailand. In E. Buffetaut, G. Cuny, J. Le Loeuff, V. Suteethorn (eds.), Late Palaeozoic and Mesozoic Ecosystems in SE Asia. The Geological Society of London, Special Publication 315:175-187.Valid Lauprasert Cuny et al.Lauprasert, Cuny, Thirakhupt, Suteethorn.Early CretaceousThailandYoungest Mesozoic crocodyliform yet known from Thailand Laganosuchus Sereno P. C. Larsson, H. C. E. 2009 Cretaceous crocodyliforms from the Sahara ZooKeys 28 1–143 10.3897/zookeys.28.325 2009ZooK...28....1S free Valid Sereno LarsonCenomanianEchkar Formation Kem Kem BedsNiger Moroccothumb|150px|center|Laganosuchus Miadanasuchus Erin L. Rasmusson Simons Gregory A. Buckley 2009 New Material of "Trematochampsa" Oblita (Crocodyliformes, Trematochampsidae) from the Late Cretaceous of Madagascar Journal of Vertebrate Paleontology 29 2 599–604 10.1671/039.029.0224 2009JVPal..29..599R 140171041 Valid Simons BuckleyCampanianMaevarano FormationMadagascarA new genus for "Trematochampsa" oblita (Buffetaut & Taquet, 1979) MorrinhosuchusIori F.V. Carvalho, I.S. 2009 Morrinhosuchus luziae, um novo Crocodylomorpha Notosuchia da Bacia Bauru, Brasil Revista Brasileira de Geociências 39 4 717–725 http://ojs.c3sl.ufpr.br/ojs2/index.php/rbg/article/viewFile/12839/11539 7 August 2010 10.25249/0375-7536.2009394717725 4 September 2025 https://web.archive.org/web/20110706162647/http://ojs.c3sl.ufpr.br/ojs2/index.php/rbg/article/viewFile/12839/11539 6 July 2011 dead free Valid Iori CarvalhoLate CretaceousAdamantina FormationBrazilNotosuchian from Brazil PenghusuchusShan, Hsi-yin Wu, Xiao-chun Cheng, Yen-nien Sato, Tamaki 2009 A new tomistomine (Crocodylia) from the Miocene of Taiwan Canadian Journal of Earth Sciences 46 7 529–555 10.1139/E09-036 http://rparticle.web-p.cisti.nrc.ca/rparticle/AbstractTemplateServlet?calyLang=eng&journal=cjes&volume=46&year=2009&issue=7&msno=e09-036 2009CaJES..46..529S 15 September 2010 subscription Valid Shan Wu et al.Shan, Wu, Cheng, Sato.Late MioceneTaiwanA tomistomine crocodilian. PolonosuchusBrusatte Stephen L. Butler Richard J. Sulej Tomasz Niedźwiedzki Grzegorz Brusatte, Stephen L. Butler, Richard J. Sulej, Tomasz Niedźwiedzki, Grzegorz 2009 The taxonomy and anatomy of rauisuchian archosaurs from the Late Triassic of Germany and Poland Acta Palaeontologica Polonica 54 2 221–230 10.4202/app.2008.0065 54044937 https://app.pan.pl/archive/published/app54/app54-221.pdf 13 August 2010 Valid Brusatte Butler et al.Brusatte, Butler, Sulej, Niedźwiedzki.Late CarnianPolandA new genus for "Teratosaurus" silesiacus (Sulej, 2005)thumb|150px|center|Polonosuchus Yacarerani Novas F. E. Pais, D. F. Pol, D. Carvalho, I. D. S. Mones, A. Scanferla, A. Riglos, M. S. 2009 Bizarre notosuchian crocodyliform with associated eggs from the Upper Cretaceous of Bolivia Journal of Vertebrate Paleontology 29 4 1316–1320 10.1671/039.029.0409 2009JVPal..29.1316N 11336/54001 140589478 free Valid Novas Pais et al.Novas, Pais, Pol, Carvalho, Mones, Scanferla, Riglos.Late Cretaceous Cajones FormationBoliviathumb|150px|center|Yacarerani Non-avian dinosaurs Research A new study on theropod furculae is published. Nesbitt S.J. Turner A.H. Spaulding M. Conrad J.L. Norell M.A. 2009 The theropod furcula Journal of Morphology 270 7 856–879 10.1002/jmor.10724 19206153 2009JMorp.270..856N 11847975 A "detailed description of the skull and mandible of the Chinese cerapodan ornithischian dinosaur Jeholosaurus shangyuanensis" is published. Barrett P.M. Han F.-L. 2009 Cranial anatomy of Jeholosaurus shangyuanensis (Dinosauria: Ornithischia) from the Early Cretaceous of China Zootaxa 2072 31–55 10.11646/zootaxa.2072.1.2 Knoll F. Padian K. de Ricqles A. 2009 Ontogenetic change and adult body size of the early ornithischian dinosaur Lesothosaurus diagnosticus: Implications for basal ornithischian taxonomy Gondwana Research 17 171–179 10.1016/j.gr.2009.03.010 Matthews J. C. Brusatte S. L. Williams S. A. Henderson M. D. 2009 The first Triceratops bonebed and its implications for gregarious behavior Journal of Vertebrate Paleontology 29 1 286–290 10.1080/02724634.2009.10010382 2009JVPal..29..286M 196608646 Williamson T. E. Carr T. D. Williams S. A. Tremaine K. 2009 Early ontogeny of pachycephalosaurine squamosals as revealed by juvenile specimens from the Hell Creek Formation, eastern Montana Journal of Vertebrate Paleontology 29 1 291–294 10.1671/039.029.0111 2009JVPal..29..291W 84516840 Bittencourt J.S. Kellner A.W.A. 2009 The anatomy and phylogenetic position of the Triassic dinosaur Staurikosaurus pricei Colbert, 1970 Zootaxa 2079 1–56 10.11646/zootaxa.2079.1.1 90574494 Chin K. Hartman J.H. Roth B. 2009 Opportunistic exploitation of dinosaur dung: fossil snails in coprolites from the Upper Cretaceous Two Medicine Formation of Montana Lethaia 42 2 185–198 10.1111/j.1502-3931.2008.00131.x 2009Letha..42..185C Maidment S.C.R. Porro L.B. 2009 Homology of the palpebral and origin of supraorbital ossifications in ornithischian dinosaurs Lethaia 43 95–111 10.1111/j.1502-3931.2009.00172.x Gates T.A. Farke A.A. 2009 Biostratigraphic and biogeographic implications of a hadrosaurid (Ornithopoda: Dinosauria) from the Upper Cretaceous Almond Formation of Wyoming, USA Cretaceous Research 30 5 1157–1163 10.1016/j.cretres.2009.05.001 2009CrRes..30.1157G Moratalla J.J. Hernán J. 2008 Los Cayos S y D: dos afloramientos con icnitas de saurópodos, terópodos y ornitópodos en el Cretácico Inferior del área de Los Cayos (Cornago, La Rioja, España) Estudios Geológicos 64 2 161–173 10.3989/egeol.08642.043 free Taylor M.P. Wedel M.J. Naish D. 2009 Head and neck posture in sauropod dinosaurs inferred from extant animals Acta Palaeontologica Polonica 54 2 213–220 10.4202/app.2009.0007 7582320 Hadrosaur chewing study 2008–2009 hadrosaur chewing study A study titled "Quantitative analysis of dental microwear in hadrosaurid dinosaurs, and the implications for hypotheses of jaw mechanics and feeding" is published by British paleontologists Mark Purnell, Paul Barrett and student Vince Williams. The paper examined the chewing methods and diet of hadrosaurid ("duck billed") dinosaurs from the Late Cretaceous period. The scientists analyzed hundreds of microscopic scratches on the teeth of a fossilized Edmontosaurus jaw, and believe they determined exactly how a hadrosaur broke down and ate its food, which had previously eluded researchers. The study found hadrosaurs had a unique way of eating unlike any creature living today. In contrast to a flexible lower jaw joint prevalent in today's mammals, hadrosaurs had a unique hinge between the upper jaws and the rest of its skull. The team found the dinosaur's upper jaws pushed outwards and sideways while chewing, as the lower jaw slid against the upper teeth. The study also concluded that hadrosaurs likely grazed on horsetails and vegetation close to the ground, rather than browsing higher-growing leaves and twigs. However, Purnell said these conclusions were less secure than the more conclusive evidence regarding the motion of teeth while chewing. Previous studies found contradictory conclusions, and the issue remains a subject of debate. The findings were published on June 30, 2009, in the journal, The Proceedings of the National Academy of Sciences. Purnell said no previous study had ever employed this method of analyzing microscopic teeth scratches, and that the method could be used to study other areas of scientific research. New taxa Data courtesy of George Olshevky's dinosaur genera list. Olshevsky George Dinogeorge's Dinosaur Genera List http://www.polychora.com/dinolist.html 2009-01-05 2011-07-15 https://web.archive.org/web/20110715102453/http://www.polychora.com/dinolist.html dead ~44 dinosaur genera were erected in 2009. Name Status Authors Discovery year Age Unit Location Notes Images AdeopapposaurusMartínez Ricardo N. 2009 Adeopapposaurus mognai, gen. et sp. nov (Dinosauria: Sauropodomorpha), with comments on adaptations of basal sauropodomorpha Journal of Vertebrate Paleontology 29 1 142–164 10.1671/039.029.0102 2009JVPal..29..142M 85074392 Valid Ricardo N. MartínezCañón del Colorado Formation Argentinathumb|150px|center|Adeopapposaurus AerosteonSereno PC, Martinez RN, Wilson JA, Varricchio DJ, Alcober OA Sep 2008 Evidence for Avian Intrathoracic Air Sacs in a New Predatory Dinosaur from Argentina PLOS ONE 3 9 e3303 18825273 2553519 10.1371/journal.pone.0003303 2008PLoSO...3.3303S Kemp Tom etal free Valid Sereno R. N. Martinez et al.Sereno, R. N. Martinez, J. A. Wilson, Varricchio, Alcober.Rio Colorado Formation Argentinathumb|150px|center|AerosteonAlbalophosaurus Ohashi T. Barrett P. M. 2009 A new ornithischian dinosaur from the Lower Cretaceous Kuwajima Formation of Japan Journal of Vertebrate Paleontology 29 3 748–757 10.1671/039.029.0306 2009JVPal..29..748O 128546047 Valid Ohashi BarrettKuwajima Formation Japan AlbertonykusLongrich Nicholas R. Currie, Philip J. 2009 Albertonykus borealis, a new alvarezsaur (Dinosauria: Theropoda) from the Early Maastrichtian of Alberta, Canada: Implications for the systematics and ecology of the Alvarezsauridae Cretaceous Research 30 1 239–252 10.1016/j.cretres.2008.07.005 2009CrRes..30..239L Valid Longrich Currielower MaastrichtianHorseshoe Canyon Formation Canada (Alberta)thumb|150px|center|Albertonykus AnchiornisXu, X., Zhao, Q., Norell, M., Sullivan, C., Hone, D., Erickson, G., Wang, X., Han, F. and Guo, Y. (2009). "A new feathered maniraptoran dinosaur fossil that fills a morphological gap in avian origin." Chinese Science Bulletin, six pages, accepted November 15, 2008. Valid Xu X. Zhao Q. et al.Xu X., Zhao Q., Norell, C. Sullivan, Hone, Erickson, Wang X. L., Han F., Guo.Tiaojishan Formation Chinathumb|150px|center|AnchiornisAngulomastacator Wagner J.R. Lehman T.M. 2009 An Enigmatic New Lambeosaurine Hadrosaur (Reptilia: Dinosauria) from the Upper Shale Member of the Campanian Aguja Formation of Trans-Pecos Texas Journal of Vertebrate Paleontology 29 2 605–611 10.1671/039.029.0208 2009JVPal..29..605W 128555861 Valid J. R. Wagner LehmanAguja Formation US (Texas)ArenysaurusPereda-Suberbiola Xabier José Ignacio Canudo Penélope Cruzado-Caballero José Luis Barco Nieves López-Martínez Oriol Oms José Ignacio Ruiz-Omeñaca 2009 The last hadrosaurid dinosaurs of Europe: A new lambeosaurine from the Uppermost Cretaceous of Aren (Huesca, Spain) Comptes Rendus Palevol 8 6 559–572 10.1016/j.crpv.2009.05.002 2009CRPal...8..559P https://eprints.ucm.es/10010/1/Pereda-Suberbiola%2609-Arenysaurus.pdf Valid Xabier Pereda-Suberbiolaa José Ignacio Canudob et al.Xabier Pereda-Suberbiolaa, José Ignacio Canudob, Penélope Cruzado-Caballerob, José Luis Barcoc, Nieves López-Martínezd, Oriol Omse, José Ignacio Ruiz-Omeñaca. SpainA Spanish Lambeosaurine.AustralovenatorMolnar Ralph E. Flannery, Timothy F. Rich, Thomas H.V. 1981 An allosaurid theropod dinosaur from the Early Cretaceous of Victoria, Australia Alcheringa 5 141–146 10.1080/03115518108565427 2 1981Alch....5..141M Valid Hocknull White et al.Winton Formation AustraliaAn Australian Megaraptor. Specimen named "Banjo"thumb|150px|center|Australovenator Baotianmansaurus Zhang X. Lü J. Xu L. Li J. Yang L.K. Hu W. Jia S. Ji Q. Zhang C. 2009 A new sauropod dinosaur from the Late Cretaceous Gaogou Formation of Nanyang, Henan Province Acta Geologica Sinica (English Edition) 83 2 212–221 10.1111/j.1755-6724.2009.00032.x 2009AcGlS..83..212Z 129422997 Valid Zhang, X. Lü, J. et al.Zhang, X., Lü, J., Xu, L., Li, J., Yang, L.K., Hu, W., Jia, S., Ji, Q. Zhang, C.Gaogou Formation ChinaBarrosasaurus Salgado L. Coria R. A. 2009 Barrosasaurus casamiquelai gen. et sp. nov., a new titanosaur (Dinosauria, Sauropoda) from the Anacleto Formation (Late Cretaceous: early Campanian) of Sierra Barrosa (Neuquén, Argentina) Zootaxa 2222 1–16 10.11646/zootaxa.2222.1.1 Valid Salgado CoriaAnacleto Formation Argentina Ceratonykus Alifanov V.R. Barsbold R. 2009 Ceratonykus oculatus gen. et sp. nov., a new dinosaur (?Theropoda, Alvarezsauria) from the Late Cretaceous of Mongolia" (in English) Paleontological Journal 43 1 86–99 10.1134/S0031030109010109 129589208 Valid Alifanov BarsboldBarun Goyot Formation Mongoliathumb|150px|center|Ceratonykus DiamantinasaurusHocknull Scott A. White, Matt A. Tischler, Travis R. Cook, Alex G. Calleja, Naomi D. Sloan, Trish Elliott, David A. 2009 New Mid-Cretaceous (Latest Albian) Dinosaurs from Winton, Queensland, Australia PLOS ONE 4 7 10.1371/journal.pone.0006190 e6190 19584929 2703565 2009PLoSO...4.6190H Sereno Paul free Valid Hocknull White et al.Winton Formation AustraliaAn Australian Titanosaur.thumb|150px|center|DiamantinasaurusElrhazosaurusGalton Peter M. Peter Galton 2009 Notes on Neocomian (Late Cretaceous) ornithopod dinosaurs from England – Hypsilophodon, Valdosaurus, "Camptosaurus", "Iguanodon" – and referred specimens from Romania and elsewhere Revue de Paléobiologie 28 1 211–273 http://www.ville-ge.ch/mhng/paleo/paleo-pdf/28-1/pal-28-1-11.pdf 7 January 2010 Valid GaltonElrhaz FormationNiger Helioceratops Jin L. Chen J. Zan S. Godefroit P. 2009 A new basal neoceratopsian dinosaur from the Middle Cretaceous of Jilin Province, China Acta Geologica Sinica (English Edition) 83 2 200–206 10.1111/j.1755-6724.2009.00023.x 2009AcGlS..83..200J 129656123 Valid Jin Chen et al.Jin, Chen, Zan, Godefroit.Quantou Formation Chinathumb|Helioceratops HesperonychusLongrich, N.R. and Currie, P.J. (2009). "A microraptorine (Dinosauria–Dromaeosauridae) from the Late Cretaceous of North America." Proceedings of the National Academy of Sciences, 106(13) .Valid Longrich CurrieDinosaur Park Formation CanadaSmallest known dinosaur from North America.thumb|150px|center|HesperonychusJintasaurus You H.-L. Li D.-Q. 2009 A new basal hadrosauriform dinosaur (Ornithischia: Iguanodontia) from the Early Cretaceous of northwestern China Canadian Journal of Earth Sciences 46 12 949–957 10.1139/E09-067 2009CaJES..46..949Y Li Da-Qing Valid You LiXinminpu Group China KinnareemimusBuffetaut, E., Suteethorn, V. and Tong, H. (2009). "An early 'ostrich dinosaur' (Theropoda: Ornithomimosauria) from the Early Cretaceous Sao Khua Formation of NE Thailand", pp. 229-243 IN E. Buffetaut, G. Cuny, J. Le Loeuff & V. Suteethorn (eds.), Late Palaeozoic and Mesozoic Ecosystems in SE Asia Geological Society, London, Special Publications 315: 229-243. 10.1144/SP315.16Valid Buffetaut Suteethorn TongSao Khua Formation Thailand KolA.H. Turner S.J. Nesbitt M.A. Norell A Large Alvarezsaurid from the Cretaceous of Mongolia American Museum Novitates 3648 1–14 2009 10.1206/639.1 2246/5967 59459861 https://zenodo.org/record/4712967 Valid Turner Nesbitt NorellDjadochta Formation Mongoliathumb|150px|center|KolLeshansaurusF. Li; Peng G.; Ye Y.; Jiang S.; and Huang, D. (2009). "A new carnosaur from the Late Jurassic of Qianwei, Sichuan, China". Acta Geologica Sinica 83(9): 1203–1213. Abstract https://web.archive.org/web/20201106120716/http://en.cnki.com.cn/Article_en/CJFDTotal-DZXE200909002.htm 2020-11-06 .Valid Li Peng et al.Li, Peng, Jiang, Huang.Shangshaximiao Formation China Levnesovia Hans-Dieter Sues Averianov Alexander 2009 A new basal hadrosauroid dinosaur from the Late Cretaceous of Uzbekistan and the early radiation of duck-billed dinosaurs Proceedings of the Royal Society B 276 1667 2549–2555 10.1098/rspb.2009.0229 19386651 2686654 Valid Sues AverianovBissekty Formation UzbekistanThe oldest Hadrosauroidean Limusaurus Xu X. Clark J. M. Mo J. Choiniere J. Forster C. A. Erickson G. M. Hone D. W. E. Sullivan C. Eberth D. A. Nesbitt S. Zhao Q. Hernandez R. Jia C. K. Han F. L. Guo Y. A Jurassic ceratosaur from China helps clarify avian digital homologies 10.1038/nature08124 Nature 459 7249 940–944 2009 19536256 2009Natur.459..940X 4358448 https://doc.rero.ch/record/209594/files/PAL_E4066.pdf Valid XuShishugou Formation ChinaThe first Asian ceratosaur to be discoveredthumb|150px|center|Limusaurus Luoyanggia Lü Xu L. Jiang X. Jia S. Li M. Yuan C. Zhang X. Ji Q. 2009 A preliminary report on the new dinosaurian fauna from the Cretaceous of the Ruyang Basin, Henan Province of central China Journal of the Paleontological Society of Korea 25 43–56 Valid Lü Xu et al.Lü, Xu, Jiang, Jia, Li, Yuan, Zhang, Ji.Haoling Formation ChinaAn oviraptorosaur Malarguesaurus González Riga B. J. Previtera E. Pirrone C.A. 2009 Malarguesaurus florenciae gen. et sp. nov., a new titanosauriform (Dinosauria, Sauropoda) from the Upper Cretaceous of Mendoza, Argentina Cretaceous Research 30 1 135–148 10.1016/j.cretres.2008.06.006 2009CrRes..30..135G Valid González Riga Previtera PirronePortezuelo Formation Argentina MinotaurasaurusMiles Clifford A. Miles, Clark J. 2009 Skull of Minotaurasaurus ramachandrani, a new Cretaceous ankylosaur from the Gobi Desert Current Science 96 1 65–70 http://www.ias.ac.in/currsci/jan102009/65.pdf 7 January 2010 Valid Clifford A. Miles Clark J. Mikes Mongoliathumb|150px|center|Minotaurasaurus MiragaiaMateus Octávio Maidment, Susannah C.R. Christiansen, Nicolai A. A new long-necked 'sauropod-mimic' stegosaur and the evolution of the plated dinosaurs Proceedings of the Royal Society B 2009 276 1663 1815–1821 10.1098/rspb.2008.1909 19324778 2674496 Valid Mateus Maidment ChristiansenSobral Unit PortugalLong-necked stegosaur.center|thumb|150x150px|Miragaia Nothronychus graffamiZanno L. E. Gillette D. D. Albright L. B. Titus A. L. 2009 A new North American therizinosaurid and the role of herbivory in predatory dinosaur evolution Proceedings of the Royal Society B 276 1672 3505−3511 10.1098/rspb.2009.1029 free 2817200 19605396 30244145 Valid Zanno et al.'Tropic Shale Formation USA therizinosaurid.thumb|150px|center|Nothronychus graffamiOwenodonValid GaltonPurbeck Limestone UKthumb|150px|center|Owenodon Panphagia Martinez RN Alcober OA Alcober Oscar A. 2009 A Basal Sauropodomorph (Dinosauria: Saurischia) from the Ischigualasto Formation (Triassic, Carnian) and the Early Evolution of Sauropodomorpha PLOS ONE 4 2 e4397 10.1371/journal.pone.0004397 19209223 2635939 2009PLoSO...4.4397M Sereno Paul free Valid Martinez AlcoberIschigualasto Formation ArgentinaOne of the most basal known sauropodomorph.thumb|150px|center|PanphagiaQiaowanlong Valid You LiXinminpu Group Chinathumb|150px|center|QiaowanlongRaptorexSereno P. Tan L. Brusatte S. L. Kriegstein H. J. Zhao X. Cloward K. amp 2009 Tyrannosaurid skeletal design first evolved at small body size Science 326 5951 418–422 10.1126/science.1177428 19762599 2009Sci...326..418S 1953405 Sereno Brusatte et al.Sereno, Brusatte, Kriegstein, Zhao, Cloward.Yixian Formation ChinaTyrannosauroideathumb|150px|center|Raptorex Ruyangosaurus Lu J Xu L Jia S Zhang X Zhang J Yang L You H Ji Q 2009 A new gigantic sauropod dinosaur from the Cretaceous of Ruyang, Henan, China Geological Bulletin of China 28 1 Valid Lu Xu et al.Lu, Xu, Jia, Zhang, Zhang, Yang, You, Ji.Haoling Formation Chinathumb|150px|center|Ruyangosaurus Shaochilong Brusatte S. Benson R. Chure D. Xu X. Sullivan C. Hone D. 2009 The first definitive carcharodontosaurid (Dinosauria: Theropoda) from Asia and the delayed ascent of tyrannosaurids https://www.pure.ed.ac.uk/ws/files/8232136/PDF_Brusatteetal2009FirstDefinitiveCarcharodontosaurid.pdf Naturwissenschaften 19488730 96 9 1051–1058 10.1007/s00114-009-0565-2 2009NW.....96.1051B 25532873 Valid Brusatte Benson et al.Brusatte, Benson, Chure, Xu, Sullivan, Hone.Miaogou Formation Chinathumb|150px|center|Shaochilong ShidaisaurusWu X. Currie, P.J. Dong Z. Pan S. Tang W. A New Theropod Dinosaur from the Middle Jurassic of Lufeng, Yunnan, China Acta Geologica Sinica 83 1 2009 10.1111/j.1755-6724.2009.00002.x 9–24 2009AcGlS..83....9W 140695367 Valid Wu Currie et al.Wu, Currie, Dong, Pan, Tang.Chuanjie Formation ChinaSinotyrannus Ji Q. Ji S.-A. Zhang L.-J. 2009 First large tyrannosauroid theropod from the Early Cretaceous Jehol Biota in northeastern China Geological Bulletin of China 28 10 1369–1374 Valid Ji Ji ZhangJiufotang Formation China Skorpiovenator Canale J.I. Scanferla C.A. Agnolin F. Novas F.E. 2009 New carnivorous dinosaur from the Late Cretaceous of NW Patagonia and the evolution of abelisaurid theropods Naturwissenschaften 19057888 96 3 409–414 10.1007/s00114-008-0487-4 2009NW.....96..409C 11336/52024 23619863 free Valid Canale Scanferla et al.Canale, Scanferla, Agnolin, Novas.Huincul Formation Argentinacenter|thumb|150x150px|SkorpiovenatorSpinophorosaurus Remes K. Ortega F. Fierro I. Ferrer J.M.M. Joger U. Kosma R. Ide Oumarou Amadou Maga Abdoulaye Ide O. A. Maga A A new basal sauropod dinosaur from the Middle Jurassic of Niger and the early evolution of sauropoda PLOS ONE 2009 4 9 e6924 10.1371/journal.pone.0006924 2009PLoSO...4.6924R Farke Andrew Allen 19756139 2737122 free Remes Ortega et al.Remes, Ortega, Fierro, Joger, Kosma, Ferrer.Irhazer Group NigerSauropodathumb|150px|center|SpinophorosaurusTatankacephalus Parsons W.L. Parsons K.M. 2009 A new ankylosaur (Dinosauria: Ankylosauria) from the Lower Cretaceous Cloverly Formation of central Montana Canadian Journal of Earth Sciences 46 10 721–738 10.1139/E09-045 2009CaJES..46..721S Parsons Kristen M. Valid Parsons ParsonsCloverly Formation US (Montana)TawaNesbitt S. J. Smith N. D. Irmis R. B. Turner A. H. Downs A. Norell M. A. amp 2009 A complete skeleton of a Late Triassic saurischian and the early evolution of dinosaurs Science 326 5959 1530–1533 10.1126/science.1180350 20007898 2009Sci...326.1530N 8349110 .Valid Nesbitt Smith et al.Nesbitt, Smith, Irmis, Turner, Downs, Norell.Chinle Formation US (New Mexico)alt=|center|thumb|150x150px|TawaTethyshadrosDalla Vecchia F. M. 2009 Tethyshadros insularis, a new hadrosauroid dinosaur (Ornithischia) from the Upper Cretaceous of Italy Journal of Vertebrate Paleontology 29 4 1100–1116 10.1671/039.029.0428 2009JVPal..29.1100D 198128196 Valid Dalla VecchiaLipica Formation ItalyTianyulong 10.1038/nature07856 Zheng X-.T. You H.-L. Xu X. Dong Z.-M. 2009 An Early Cretaceous heterodontosaurid dinosaur with filamentous integumentary structures Nature 458 19 333–336 19295609 2009Natur.458..333Z 4423110 Valid Zheng You et al.Zheng, You, Xu, Dong.Tiaojishan Formation Chinathumb|150px|center|TianyulongWintonotitanValid Hocknull White et al.Hocknull, White, Tischler, Cook, Calleja, Sloan, Elliott.Winton Formation Australiathumb|150px|center|Wintonotitan Xianshanosaurus Valid Lü Xu et al.Haoling Formation ChinaA sauropod"Xinghesaurus"Nomen nudum Hasegawa Carpenter et al.Hasegawa, Carpenter, Lamanna, Xu. Name published without scientific description in Japanese guidebook "Dinosaur Expo 2009: The Miracle of Deserts"ZanabazarM.A. Norell Makovicky, P.J. Bever, G.S. Balanoff, A.M. Clark, J.M. Barsbold, R. Rowe, T. A Review of the Mongolian Cretaceous Dinosaur Saurornithoides (Troodontidae: Theropoda) American Museum Novitates 3654 1–63 2009 10.1206/648.1 2246/5973 https://www.biodiversitylibrary.org/bibliography/169010 Valid Norell Makovicky et al.Norell, Makovicky, Bever, Balanoff, Clark, Barsbold, Rowe.Nemegt Formation Mongoliathumb|150px|center|Zanabazar Aves Research Anfinson O.A. Lockley M.G. Kim S.H. Kim K.S. Kim J.Y. 2009 First report of the small bird track Koreanaornis from the Cretaceous of North America: implications for avian ichnotaxonomy and paleoecology Cretaceous Research 30 4 885–894 10.1016/j.cretres.2009.02.001 2009CrRes..30..885A https://doc.rero.ch/record/16674/files/PAL_E3274.pdf Lockley M. Chin K. Houck K. Matsukawa M. Kukihara R. 2009 New interpretations of Ignotornis, the first-reported Mesozoic avian footprints: implications for the paleoecology and behavior of an enigmatic Cretaceous bird Cretaceous Research 30 4 1041–1061 10.1016/j.cretres.2009.04.001 2009CrRes..30.1041L https://doc.rero.ch/record/16726/files/PAL_E3774.pdf Bell A. Everhart M.J. 2009 A new specimen of Parahesperornis (Aves: Hesperornithiformes) from the Smoky Hill Chalk (Early Campanian) of western Kansas Transactions of the Kansas Academy of Science 112 1/2 7–14 10.1660/062.112.0202 86083098 Newly described birds Name Novelty Status Authors Age Unit Location Notes ImagesAlamitornis minutus Federico L. Agnolin Agustín G. Martinelli 2009 Fossil birds from the Late Cretaceous Los Alamitos Formation, Río Negro Province, Argentina https://www.researchgate.net/publication/233755248 Journal of South American Earth Sciences 27 1 42–49 10.1016/j.jsames.2008.09.003 2009JSAES..27...42AGen. nov. et Sp. nov.ValidFederico L. Agnolin Agustín G. MartinelliLate CretaceousLos Alamitos Formation ArgentinaPlaced in ?Patagopterygiformes by Agnolin and Martinelli.Australotadorna alecwilsoni 2009 Trevor H. Worthy Descriptions And Phylogenetic Relationships Of Two New Genera and Four New Species Of Oligo-Miocene Waterfowl (Aves: Anatidae) from Australia https://www.researchgate.net/publication/227774047 Zoological Journal of the Linnean Society 156 2 411–454 10.1111/j.1096-3642.2008.00483.x 85990677 free Gen. nov. et Sp. nov.ValidTrevor H. WorthyLate OligoceneEtadunna Formation AustraliaAn Anatidae.Bonasa nini Antonio Sánchez Marco 2009 New Iberian Galliforms and Reappraisal of Some Pliocene and Pleistocene Eurasian Taxa Journal of Vertebrate Paleontology 29 4 1148–1161 10.1671/039.029.0421 2009JVPal..29.1148M 130450799 Sp. nov.ValidAntonio Sánchez MarcoEarly PleistoceneLower Elefante, TE 13 layer SpainA Phasianidae.Calidris janossyi 2009 Jenö Kessler Új Eredmények a Kárpát-medence Neogén és Negyedidöszaki Madárvilágához II. Rész (New Results with Regard to the Neogene and Quaternary Avifauna of the Carpathian Basin Part II) https://epa.oszk.hu/01600/01635/00329/pdf/EPA01635_foldtani_kozlony_2009_139_3_251-271.pdf Földtani Közlöny 139 3 251–271 Sp. nov.ValidJenö KesslerLate MioceneMN 13 HungaryOriginally described as a member of Scolopacidae belonging to the genus Calidris; Zelenkov, Volkova and Gorobets (2016) reinterpreted it as a member of Turnicidae and transferred it to the genus Ortyxelos.Nikita V. Zelenkov Natalia V. Volkova Leonid V. Gorobets 2016 Late Miocene buttonquails (Charadriiformes, Turnicidae) from the temperate zone of Eurasia Journal of Ornithology 157 1 85–92 10.1007/s10336-015-1251-0 2016JOrni.157...85Z 14951696 Calonectris kurodai Storrs L. Olson 2009 A New Diminutive Species of Shearwater of the Genus Calonectris (Aves: Procellariidae) from the Middle Miocene Calvert Formation of Chesapeake Bay Proceedings of the Biological Society of Washington 122 4 466–470 10.2988/09-19.1 86663993 Sp. nov.ValidStorrs L. OlsonMiddle MioceneCalvert Formation US (Virginia)A Procellariidae, the smallest of the genus.Cariama santacrucensis Jorge I. Noriega Sergio F. Vizcaino Susana Bargo 2009 First Record and a New Species of Seriema (Aves: Ralliformes: Cariamidae) from Santacrucian (Early-Middle Miocene) Beds of Patagonia Journal of Vertebrate Paleontology 29 2 620–626 10.1671/039.029.0216 2009JVPal..29..620N 11336/80002 86428640 free Sp. nov.DisputedJorge I. Noriega Sergio F. Vizcaino Susana BargoEarly-Middle MioceneEstancia La Costa Member, Santa Cruz Formation ArgentinaFederico L. Agnolin, 2009, made it the type species of the separate genus Noriegavis Agnolin, 2009. Federico L. Agnolin 2009 Sistemática y Filogenia de las Aves Fororracoideas (Gruiformes, Cariamae) http://www.fundacionazara.org.ar/img/libros/sistematica-y-filogenia-de-aves-fororracoideas.pdf Fundación de Historia Natural Felix de Azara 1–79 2015-04-16 2018-10-01 https://web.archive.org/web/20181001034056/http://www.fundacionazara.org.ar/img/libros/sistematica-y-filogenia-de-aves-fororracoideas.pdf dead Originally interpreted as a seriema; Noriega & Mayr (2017) reinterpreted it as a member of the falconid genus Thegornis of uncertain specific assignment, on the basis of a reexamination of the holotype specimen.Jorge I. Noriega Gerald Mayr 2017 The systematic affinities of the putative seriema Noriegavis santacrucensis (Noriega et al., 2009) from the Miocene of Argentina Contribuciones del Museo Argentino de Ciencias Naturales 7 133–139 Charadrius lambrechti.Sp. nov.ValidJenö KesslerLate MioceneMN 13 HungaryA Scolopacidae.Clangula matraensis 2009 Jenö Kessler Új Eredmények a Kárpát-medence Neogén és Negyedidöszaki Madárvilágához I. Rész (New Results with Regard to the Neogene and Quaternary Avifauna of the Carpathian Basin Part I) https://www.matarka.hu/cikk_list.php?fusz=43615 Földtani Közlöny 139 1 67–82 Sp. nov.ValidJenö KesslerMiddle MioceneMN 6–8 HungaryAn Anatidae.Confuciusornis feducciai Zhang Zuhui Gao Chunling Meng Qingjin Liu Jinyuan Hou Lianhai Zheng Guangmei 2009 Diversification in an Early Cretaceous Avian Genus: Evidence from a New Species of Confuciusornis from China Journal of Ornithology 150 4 783–790 10.1007/s10336-009-0399-x 2009JOrni.150..783Z 21418230 Sp. nov.DisputedZuhui Zhang Chunling Gao Qingjin Meng Jinyuan Liu Lianhai Hou Guangmei ZhengEarly CretaceousYixian Formation ChinaA member of the family Confuciusornithidae. Considered to be a junior synonym of Confuciusornis sanctus by Wang, O'Connor & Zhou (2018).Min Wang Jingmai O'Connor Zhong-He Zhou 2018 A taxonomical revision of the Confuciusornithiformes (Aves: Pygostylia) Vertebrata PalAsiatica in press 10.19615/j.cnki.1000-3118.180530 Cygnopterus neogradiensis Jenö Kessler János Hír 2009 A New Anserid Species from the Neogene of Hungary Fragmenta Palaeontologica Hungarica 27 2 97–101 Sp. nov.ValidJenö Kessler János HírMiddle MioceneSajóvölgy Formation, MN 7–8 HungaryAn Anatidae.Diomedeoides harmati Jenö Kessler 2009 The Oldest Modern Bird (Ornithurinae) Remains from the Early Oligocene of Hungary Fragmenta Palaeontologica Hungarica 27 2 93–96 Sp. nov.ValidJenö KesslerEarly OligoceneMP 24 HungaryA Diomedeoididae Fischer, 1985, placed in Rupelornis van Beneden, 1871 by Gerald Mayr and Thierry Smith, 2012Egretta polgardiensis Sp. nov.DisputedJenö KesslerLate MioceneMN 13 HungaryOriginally described as a member of Ardeidae and a species of Egretta; however, Zelenkov (2017) considered this species to be a junior synonym of the barn-owl species Tyto campiterrae Jánossy (1991).Elbretornis bonapartei 2009 Euenantiornithine birds from the Late Cretaceous of El Brete (Argentina) http://www.ria.ie/RIA/files/40/40a4c454-eef2-4de0-922d-48ae0c42037f.pdf Irish Journal of Earth Sciences 27 15–62 10.3318/ijes.2010.27.15 Cyril A. Walker Gareth J. Dyke 129573066 dead https://web.archive.org/web/20120320093759/http://www.ria.ie/RIA/files/40/40a4c454-eef2-4de0-922d-48ae0c42037f.pdf 2012-03-20 Gen. nov. et Sp. nov.ValidCyril A. Walker Gareth J. DykeMaastrichtianLecho Formation ArgentinaAn Enantiornithes Walker, 1981, Euenantiornithes Chiappe, 2002, this is the type species of the new genus.Eoanseranas handae Trevor H. Worthy John D. Scanlon 2009 An Oligo-Miocene Magpie Goose (Aves: Anseranatidae) from Riversleigh, Northwestern Queensland, Australia https://www.researchgate.net/publication/232677982 Journal of Vertebrate Paleontology 29 1 205–211 10.1671/039.029.0103 2009JVPal..29..205W 129930435 Gen. nov. et Sp. nov.ValidTrevor H. Worthy John D. ScanlonLate Oligocene – Early MioceneRiverleigh World Heritage Property AustraliaAn Anseranatidae, this is the type species of the new genus.Gansugyps linxiaensis Zihui Zhang Xiaoting Zheng Guangmei Zheng Lianhai Hou 2009 A New Old World Vulture (Falconiformes: Accipitridae) from the Miocene of Gansu Province, North Western China http://gsw.lyu.edu.cn/picture/article/88/47/7a/7a70010f4bcfb325fea952fd7c1c/19e4f8d9-4683-4369-8bfe-c08427fd260e.pdf Journal of Ornithology 151 1 401–408 10.1007/s10336-009-0468-1 37386553 2015-04-16 https://web.archive.org/web/20141221135801/http://gsw.lyu.edu.cn/picture/article/88/47/7a/7a70010f4bcfb325fea952fd7c1c/19e4f8d9-4683-4369-8bfe-c08427fd260e.pdf 2014-12-21 dead Gen. nov. et Sp. nov.ValidZhang Zihui Zheng Xiaoting Zheng Guangmei Hou LianhaiLate MioceneUpper Liushu Formation ChinaAn Accipitridae, this is the type species of the new genus.Heliadornis minor Sp. nov.ValidJenö KesslerLate PlioceneMN 15–16 SlovakiaA Phaethontidae.Heliornis sumeghensis Sp. nov.ValidJenö KesslerLate MioceneMN 11–12 HungaryA HeliornithidaeJianchangornis microdonta Zhou Zhonghe Zhang Fucheng Li Zhiheng 2009 A new basal ornithurine bird (Jianchangornis microdonta gen. et sp. nov.), from the Lower Cretaceous of China http://www.ivpp.cas.cn/cbw/gjzdwxb/xbwzxz/200911/P020091104363560998049.pdf Vertebrata PalAsiatica 47 4 299–310 10.19615/j.cnki.1000-3118.2009.04.004 Gen. nov. et Sp. nov.ValidZhou Zhonghe Zhang Fucheng Li ZhihengEarly Cretaceous AlbianJiufotang Formation ChinaA basal Ornithurae Haeckel, 1866, this is the type species of the new genus.Martinavis minor Sp. nov.ValidCyril A. Walker Gareth J. DykeLate Cretaceous MaastrichtianLecho Formation ArgentinaAn Enantiornithes Walker, 1981, Euenantiornithes Chiappe, 2002.Martinavis saltariensis Sp. nov.ValidCyril A. Walker Gareth J. DykeLate Cretaceous MaastrichtianLecho Formation ArgentinaAn Enantiornithes Walker, 1981, Euenantiornithes Chiappe, 2002.Martinavis whetstonei Sp. nov.ValidCyril A. Walker Gareth J. DykeLate Cretaceous MaastrichtianLecho Formation ArgentinaAn Enantiornithes Walker, 1981, Euenantiornithes Chiappe, 2002.Megalocoturnix cordoni Gen. nov. et Sp. nov.ValidAntonio Sánchez MarcoEarly PlioceneRuscinian, MN 15 SpainA Phasianidae, this is the type species of the new genus.Megapaloelodus peiranoi Federico L. Agnolin 2009 Una nueva especie del género Megapaloelodus (Aves: Phoenicopteridae: Palaelodinae) del Mioceno Superior del noroeste de Argentina http://www.scielo.org.ar/pdf/rmacn/v11n1/v11n1a04.pdf Revista del Museo Argentino de Ciencias Naturales Nueva Series 11 1 23–32 10.22179/REVMACN.11.267Sp. nov.ValidFederico L. AgnolinLate MioceneAndalhualá Formation ArgentinaA Phoenicopteriformes, Palaelodidae Stejneger, 1885.Mergus minor Sp. nov.ValidJenö KesslerMiddle MioceneMN 6–8 HungaryAn Anatidae.Miocepphus blowi Erik Wijnker Storrs L. Olson 2009 A Revision of the Fossil Genus Miocepphus and Other Miocene Alcidae (Aves: Charadriiformes) of the Western North Atlantic Ocean https://www.researchgate.net/publication/224922716 Journal of Systematic Palaeontology 7 4 471–487 10.1017/s1477201909990022 2009JSPal...7..471W 53352434 Gen. nov. et Sp. nov.ValidErik Wijnker Storrs L. OlsonLate MioceneEastover Formation; Breda Formation Mill US (Virginia) NetherlandsAn Alcidae.Miocepphus bohaskai Sp. nov.ValidErik Wijnker Storrs L. OlsonLate MioceneEastover Formation; Breda Formation Mill US (Maryland and Virginia) NetherlandsAn Alcidae.Miocepphus mergulellus Sp. nov.ValidErik Wijnker Storrs L. OlsonLate MioceneEastover Formation US (North Carolina)An Alcidae.''[[Palaeocryptonyx novaki]] |Sp. nov. | Valid | Antonio Sánchez Marco | Late Pliocene | Villanyan, MN 16 | Spain | A Phasianidae. | |- |Paracoracias occidentalis Julia A. Clarke Daniel T. Ksepka N. Adam Smith Mark A. Norell 2009 Combined Phylogenetic Analysis of a New North American Fossil Species Confirms Widespread Eocene Distribution for Stem Rollers (Aves, Coracii) Zoological Journal of the Linnean Society 157 3 586–611 10.1111/zoj.12181 |Gen. nov. et Sp. nov. | Valid | Julia A. Clarke Daniel T. Ksepka N. Adam Smith Mark A. Norell | Eocene | Green River Formation | US (Wyoming) | A stem Coraiiformes. | |- |Pinpanetta fromensis |Sp. nov. | Valid | Trevor H. Worthy | Late Oligocene | Etadunna Formation | Australia | An Anatidae. | |- |Pinpanetta tedfordi |Gen. nov. et Sp. nov. | Valid | Trevor H. Worthy | Late Oligocene | Etadunna Formation | Australia | An Anatidae, this is the type species of the new genus. | |- |Pinpanetta vickersrichae |Sp. nov. | Valid | Trevor H. Worthy | Late Oligocene | Etadunna Formation | Australia | An Anatidae. | |- |Podiceps csarnotatus |Sp. nov. | Valid | Jenö Kessler | Late Pliocene | MN 15-16 | Hungary | A Podicipedidae. | |- |Porzana kretzoii |Sp. nov. | Valid | Jenö Kessler | Late Miocene | MN 13 | Hungary | A member of the family Rallidae. Originally described as a species Porzana; Zelenkov (2017) transferred this species to the genus Zapornia.N. V. Zelenkov 2017 Revision of non-passeriform birds from Polgárdi (Hungary, Upper Miocene): 3. Neoaves Paleontological Journal 51 2 203–213 10.1134/S0031030117020162 2017PalJ...51..203Z 90314539 | |- |Porzana matraensis |Sp. nov. | Valid | Jenö Kessler | Middle Miocene | MN 6-8 | Hungary | A Rallidae. | |- |Primozygodactylus eunjooae Gerald Mayr Nikita V. Zelenkov 2009 New Specimens of Zygodactylid Birds from the Middle Eocene of Messel, with Description of a New Species of Primozygodactylus https://www.app.pan.pl/archive/published/app54/app54-015.pdf Acta Palaeontologica Polonica 54 1 15–20 10.4202/app.2009.b103 55370152 |Sp. nov. | Valid | Gerald Mayr Nikita V. Zelenkov | Middle Eocene | MP 11 | Germany | A Zygodactylidae Brodkorb, 1971. | |- |Pseudocepphus teres |Gen. nov. et Sp. nov. | Valid | Erik Wijnker Storrs L. Olson | Late Miocene | St. Marys Formation | US (Maryland) | An Alcidae, this is the type species of the new genus. | |- |Rapaxavis pani Eric M. Morschhauser David J. Varricchio Gao Chunling Liu Jinyuan Wang Xuri Cheng Xiadong Meng Qingjin 2009 Anatomy of the Early Cretaceous bird Rapaxavis pani, a new species from Liaoning Province, China Journal of Vertebrate Paleontology 29 2 545–554 10.1671/039.029.0210 2009JVPal..29..545M 84643293 |Gen. nov. et Sp. nov. | Valid | Eric M. Morschhauser David J. Varricchio Gao Chunling Liu Jinyuan Wang Xuri Cheng Xiadong Meng Qingjin | Early Cretaceous | Jiufotang Formation | China | An Enantiornithes Walker, 1981, Longipterygidae Zhang, Zhou, Hou et Gu, 2000, this is the type species of the new genus. | |- |Rupephaps taketake Trevor H. Worthy Suzanne J. Hand Jennifer P. Worthy Alan J. D. Tennyson R. Paul Scofield 2009 A Large Fruit Pigeon (Columbidae) from the Early Miocene of New Zealand https://www.researchgate.net/publication/227943612 The Auk 126 3 649–656 10.1525/auk.2009.08244 86799657 free |Gen. nov. et Sp. nov. | Valid | Trevor H. Worthy Suzanne J. Hand Jennifer P. Worthy Alan J. D. Tennyson R. Paul Scofield | Early Miocene, Altonian | Bannockburn Formation | New Zealand | A Columbidae, this is the type species of the new genus. | |- |Sapeornis angustis Pauline Provini Zhonghe Zhou Fucheng Zhang 2009 A New Species of the Basal Bird Sapeornis from the Early Cretaceous of Liaoning, China http://www.ivpp.cas.cn/cbw/gjzdwxb/xbwzxz/200909/P020090917534252976353.pdf Vertebrata PalAsiatica 47 3 194–207 10.19615/j.cnki.1000-3118.2009.03.003 |Sp. nov. | Valid | Pauline Provini Zhou Zhonghe Zhang Fucheng | Early Cretaceous, Aptian | Jiufotang Formation | China | A Sapeornithidae Zhou et Zhang, 2006. | |- |Selenornis steendorpensis Gerald Mayr 2009 A Small Loon and a New Species of Large Owl from the Rupelian of Belgium (Aves: Gaviiformes, Strigiformes) http://www.senckenberg.de/files/content/forschung/abteilung/terrzool/ornithologie/belgium_owl.pdf Paläontologische Zeitschrift 83 2 247–254 10.1007/s12542-009-0019-3 2009PalZ...83..247M 66209247 |Sp. nov. | Valid | Gerald Mayr | Early Oligocene, MP ?23-24 | Boom Formation | Belgium | A Tytonidae, Selenornithinae Mourer-Chauviré, 1987. | |- |Shanweiniao cooperorum Jingmai K. O'Connor Xuri Wang Luis M. Chiappe Chunling Gao Qingjin Meng Xiaodong Cheng Jinyuan Liu 2009 Phylogenetic Support for a Specialized Clade of Cretaceous Enantiornithine Birds with Information from a New Species http://geology.usc.edu/~jingmai/oconnoretal2009shanweiniao.pdf Journal of Vertebrate Paleontology 29 1 188–204 10.1080/02724634.2009.10010371 2009JVPal..29..188O 196607241 |Gen. nov. et Sp. nov. | Valid | Jingmai K. O'Connor Xuri Wang Luis M. Chiappe Chunling Gao Qingjin Meng Xiaodong Cheng Jinyuan Liu | Early Cretaceous | Yixian Formation | China | An Enantiornithes Walker, 1981, Familia Longipterygidae Zhang, Zhou, Hou et Gu, 2000, this is the type species of the new genus. | thumb|150px|center|Shanweiniao cooperorum | |- |Talpanas lippa Iwaniuk, A. L. Olson, S. L. James, H. F. 24 November 2009 Extraordinary cranial specialization in a new genus of extinct duck (Aves: Anseriformes) from Kauai, Hawaiian Islands Zootaxa Magnolia Press Auckland, New Zealand 2296 47–67 https://www.mapress.com/zootaxa/2009/f/zt02296p067.pdf 10.11646/zootaxa.2296.1.3 4176091 |Gen. nov. et Sp. nov. | Valid | Storrs L. Olson Helen F. James | Holocene | Kauai | US (Hawaii) | An Anatidae, Kaua'i Mole Duck, a Hawaiian duck that probably lived a kiwi-like lifestyle, this is the type species of the new genus. | |- |Tologuica aurorae Nikita V. Zelenkov Evgeny N. Kurochkin 2009 Neogene Phasianids (Aves: Phasianidae) of Central Asia: 1. Genus Tologuica gen. nov. https://www.academia.edu/attachments/15847058/download_file?st=MTQwODY1MDk4MCw0Ni4xMjkuNTcuMTU0LDQ1MDUyNg%3D%3D&s=sidebar Paleontological Journal 43 2 208–215 10.1134/s0031030109020130 2009PalJ...43..208Z 84965715 |Gen. nov. et Sp. nov. | Valid | Nikita V. Zelenkov Evgeny N. Kurochkin | Middle Miocene | Ööshin Formation | Mongolia | A Phasianidae, this is the type species of the new genus. | |- |Tologuica karhui |Sp. nov''.ValidNikita V. Zelenkov Evgeny N. KurochkinMiddle MioceneÖöshin Formation MongoliaA Phasianidae. Pterosaurs Lü J. 2009 A new non-pterodactyloid pterosaur from Qinglong County, Hebei Province of China Acta Geologica Sinica (English Edition) 83 2 189–199 10.1111/j.1755-6724.2009.00062.x 2009AcGlS..83..189L 247739978 Vullo R. Neraudeau D. 2009 Pterosaur remains from the Cenomanian (Late Cretaceous) Paralic Deposits of Charentes, Western France Journal of Vertebrate Paleontology 29 1 277–282 10.1080/02724634.2009.10010380 2009JVPal..29..277V 197536660 Newly named pterosaurs Name Status Authors Age Unit Location Notes ImagesCarniadactylusDalla Vecchia Fabio M. 2009 Anatomy and systematics of the pterosaur Carniadactylus (gen. n.) rosenfeldi (Dalla Vecchia, 1995) Rivista Italiana de Paleontologia e Stratigrafia 115 2 159–188Valid Dalla VecchiaLate Triassic ItalyA campylognathoidid with a wingspan of about 70 cm.Changchengopterus 10.1111/j.1755-6724.2009.00062.x Lü, J. 2009 A new non-pterodactyloid pterosaur from Qinglong County, Hebei Province of China Acta Geologica Sinica (English Edition) 83 2 189–199 2009AcGlS..83..189L 247739978 Valid LüMiddle JurassicTiaojishan Formation ChinaA primitive long-tailed pterosaur related to Dorygnathus.Ningchengopterus 10.1111/j.1755-6724.2009.00001.x Lü J. 2009 A baby pterodactyloid pterosaur from the Yixian Formation of Ningcheng, Inner Mongolia, China Acta Geologica Sinica 83 1 1–8 2009AcGlS..83....1L 85074573 Valid LüEarly CretaceousYixian Formation ChinaA pterodactyloid known from a juvenile specimen whose fossils preserved details of the flight membrane and fur.thumb|center|150px|Ningchengopterus.WukongopterusWang X. Kellner, A. W. A. Jiang S. Meng X. 2009 An unusual long-tailed pterosaur with elongated neck from western Liaoning of China Anais da Academia Brasileira de Ciências 81 4 793–812 10.1590/S0001-37652009000400016 19893903 free Valid Wang Kellner Jiang MengLate JurassicDaohugou Beds ChinaA primitive long tailed pterosaur. The type specimen shows evidence of the animal having broken its shin while alive.thumb|Wukongopterus See also Paleontology History of science Dinosaurs 2010 in archosaur paleontology 2011 in archosaur paleontology 2012 in archosaur paleontology 2013 in archosaur paleontology Notes CAL References 30em Category:2009 in paleontology