2012 in paleontology

none 2012 paleontology science paleobotany arthropod paleontology paleoentomology paleomalacology reptile paleontology archosaur paleontology mammal paleontology paleoichthyology Paleontology History of science Dinosaurs 2012 Note: In 2012, the International Code of Zoological Nomenclature was amended, with new regulations allowing the publication of new names and nomenclatural acts in zoology after 2011, in works "produced in an edition containing simultaneously obtainable copies by a method that assures [...] widely accessible electronic copies with fixed content and layout", provided that the work is registered in ZooBank before it is published, the work itself states the date of publication with evidence that registration has occurred, and the ZooBank registration states both the name of an electronic archive intended to preserve the work and the ISSN or ISBN associated with the work.International Commission on Zoological Nomenclature 2012 Amendment of Articles 8, 9, 10, 21 and 78 of the International Code of Zoological Nomenclature to expand and refine methods of publication Zootaxa 3450 1–7 http://www.mapress.com/zootaxa/2012/f/zt03450p007.pdf 10.11646/zootaxa.3450.1.1 Zhi-Qiang Zhang 2012 A new era in zoological nomenclature and taxonomy: ICZN accepts e-publication and launches ZooBank Zootaxa 3450 8 http://www.mapress.com/zootaxa/2012/f/zt03450p008.pdf 10.11646/zootaxa.3450.1.2 New scientific names appearing in electronic works are not required to be registered in ZooBank, only the works themselves are. Works containing descriptions of some of the taxa listed below were not printed on paper in 2012; however, the taxa that were described in works which were registered in ZooBank in 2012 are listed as valid. Plants 2012 in paleobotany Arthropods 2012 in arthropod paleontology The following is a summary of the arthropods described in 2012 41 arachnids 105 crustaceans 362 insects 5 merostomatans 51 trilobites 9 other arthropods Bryozoans Name Novelty Status Authors Age Unit Location Notes ImagesAcupiporaR. V. Gorjunova O. B. Weiss 2012 A new genus Acupipora gen. nov. from the Upper Carboniferous of the East European Platform and problem of classification of bryozoans of the order Fenestellida Paleontological Journal 46 1 16–28 10.1134/S0031030112010054 2012PalJ...46...16G 89097921 Gen. et comb. novValidGorjunova & WeissLate CarboniferousRussiaA member of Fenestellida. The type species is "Polypora" subborealis Schulga-Nesterenko (1951).AdmirandoporaYa. Ariunchimeg 2012 Admirandopora nom. nov., a new replacement generic name for a Carboniferous bryozoan Paleontological Journal 46 3 330 10.1134/S0031030112030045 2012PalJ...46..330A 85811901 free Nom. nov.ValidAriunchimegLower CarboniferousMongoliaA replacement name for the genus Admiranda Ariunchimeg, 1996.AntoniettellaE. Di Martino P.D. Taylor 2012 Systematics and life history of Antoniettella exigua, a new genus and species of cribrimorph bryozoan from the Miocene of East Kalimantan (Indonesia) Bollettino della Società Paleontologica Italiana 51 2 99–108 http://paleoitalia.org/archives/bollettino-spi/66/vol-51-2-2012/ 2013-01-28 2014-01-01 https://web.archive.org/web/20140101012306/http://paleoitalia.org/archives/bollettino-spi/66/vol-51-2-2012/ usurped Gen. et sp. novValidDi Martino & TaylorEarly or middle MioceneIndonesiaA cribrimorph ascophoran. The type species is Antoniettella exigua.ArgentinodictyaAndrej Ernst Marcelo Carrera 2012 Upper Ordovician (Sandbian) Bryozoan Fauna from Argentine Precordillera Journal of Paleontology 86 5 721–752 10.1666/12-024.1 2012JPal...86..721E 11336/52220 86306266 free Gen. et sp. novValidErnst & CarreraLate Ordovician (Sandbian)ArgentinaA cryptostome bryozoan, a member of Ptilodictyina. The type species is Argentinodictya lenticulata.Bigeyina iberaAndrej Ernst 2012 Fenestrate bryozoan fauna from the Lower – Middle Devonian of NW Spain Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen 264 3 205–247 10.1127/0077-7749/2012/0237 2012NJGPA.264..205E Sp. nov.ValidErnstDevonianSpainA member of Fenestrata.Celleporina medoborensisUrszula Hara Marek Jasionowski 2012 The Early Sarmatian bryozoan Celleporina medoborensis sp. nov. from the Medobory reefs of western Ukraine (Central Paratethys) Geological Quarterly 56 4 895–906 http://gq.pgi.gov.pl/gq/article/view/7861 10.7306/gq.1064 4 July 2025 2012GeolQ..56..895H free Sp. novValidHara & JasionowskiMiocene (early Sarmatian)UkraineA celleporid cheilostome bryozoan, a species of Celleporina.Chasmatopora rossaeSp. novValidErnst & CarreraLate Ordovician (Sandbian)ArgentinaA phyloporinine, a species of Chasmatopora.Dissotrypa punctataSp. nov.ValidErnstDevonianSpainA member of Fenestrata.Dissotrypa robustaSp. nov.ValidErnstDevonianSpainA member of Fenestrata.Eosemicoscinium serratumSp. nov.ValidErnstDevonianSpainA member of Fenestrata.Hemitrypa cantabricaSp. nov.ValidErnstDevonianSpainA member of Fenestrata.IberofenestellaGen. et sp. nov.ValidErnstDevonianSpainA member of Fenestrata. Genus includes new species I. wolfae.LunostomaAndrej Ernst Paul D. Taylor Jan Bohatý Patrick N. Wyse Jackson 2012 Homeomorphy in Lunostoma, a new Middle Devonian cryptostome bryozoan Paläontologische Zeitschrift 86 2 135–145 10.1007/s12542-011-0127-8 2012PalZ...86..135E 129384447 Gen. et sp. novValidErnst, Taylor, Bohatý & Wyse JacksonMiddle DevonianGermanyA cryptostome bryozoan. The type species is Lunostoma pulchra.Parvohallopora parvulaSp. novValidErnst & CarreraLate Ordovician (Sandbian)ArgentinaA trepostome bryozoan, a species of Parvohallopora.Paucipora akishinensisSp. novValidGorjunova & WeissLate Carboniferous (Moscovian)Domodedovo FormationRussiaA member of Fenestellida.Prolixicella iberaSp. nov.ValidErnstDevonianSpainA member of Fenestrata.Prolixicella parvaSp. nov.ValidErnstDevonianSpainA member of Fenestrata.Ptilodictya intermediaSp. novValidErnst & CarreraLate Ordovician (Sandbian)ArgentinaA cryptostome bryozoan, a species of Ptilodictya.PyrisinellaEmanuela Di Martino Paul D. Taylor 86922396 2012 Pyrisinellidae, a new family of anascan cheilostome bryozoans Zootaxa 3534 1–20 10.11646/zootaxa.3534.1.1 Gen. et comb. novValidDi Martino & TaylorLate Cretaceous (Maastrichtian)Peedee Formation Prairie Bluff Chalk Ripley FormationUnited StatesA member of Cheilostomata belonging to the family Pyrisinellidae. The type species is "Setosinella" meniscacantha Taylor & McKinney (2006).Rectifenestella elegantulaSp. nov.ValidErnstDevonianSpainA member of Fenestrata.Rectifenestella villayandrensisSp. nov.ValidErnstDevonianSpainA member of Fenestrata.RoryporaPaul D. Taylor 2012 A new bryozoan genus from the Jurassic of Switzerland, with a review of the cribrate colony-form in bryozoans Swiss Journal of Palaeontology 131 2 201–210 10.1007/s13358-011-0027-2 2012SwJP..131..201T 84486778 free Gen. et comb. nov.ValidTaylorJurassicSwitzerlandA new genus for "Diastopora" retiformis.Setosinella perfluxaSp. novValidDi Martino & TaylorMiocene (Langhian)IndonesiaA member of Cheilostomata belonging to the family Pyrisinellidae.SpinisinellaGen. et sp. novValidDi Martino & TaylorLate Cretaceous (Cenomanian)Czech Republic GermanySilviu O. Martha Birgit Niebuhr Joachim Scholz 2016 Kreide-Fossilien in Sachsen, Teil 2. 9. Cheilostome Bryozoen Geologica Saxonica 62 11–52 http://www.senckenberg.de/files/content/forschung/publikationen/geologicasaxonica/62/09__geologica-saxonica_62_2016_martha-et-al.pdf 2017-07-06 https://web.archive.org/web/20170923002256/http://www.senckenberg.de/files/content/forschung/publikationen/geologicasaxonica/62/09__geologica-saxonica_62_2016_martha-et-al.pdf 2017-09-23 A member of Cheilostomata belonging to the family Pyrisinellidae. The type species is S. zagorseki.Trigonodictya parvulaSp. novValidErnst & CarreraLate Ordovician (Sandbian)ArgentinaA cryptostome bryozoan, a species of Trigonodictya. Brachiopods Name Novelty Status Authors Age Unit Location Notes ImagesAmydroptychus markowitziHoward R. Feldman Mena Schemm-Gregory Fayez Ahmad Mark A. Wilson 2012 Jurassic rhynchonellide brachiopods from the Jordan Valley Acta Palaeontologica Polonica 57 1 191–204 http://app.pan.pl/article/item/app20100092.html 10.4202/app.2010.0092 2012AcPaP..57..191F 56085342 free sp novValidFeldman et al.CallovianMughanniyya FormationJordanA tetrarhynchiid rhynchonellide, a species of Amydroptychus.Anisopleurella tricostataChristian M. Ø. Rasmussen David A. T. Harper Robert B. Blodgett 2012 Late Ordovician brachiopods from West-central Alaska: systematics, ecology and palaeobiogeography Wiley-Blackwell Fossils and Strata 58 1–103 978-1-118-38417-6 10.1111/j.1502-3931.2011.00298.x https://www.academia.edu/31155178 Sp. novValidRasmussen, Harper & BlodgettLate OrdovicianFarewell TerraneUnited StatesA member of Strophomenida belonging to the group Plectambonitoidea and the family Sowerbyellidae.Bolilaspirifer riccardiiSusana E. Damborenea Miguel O. Manceñido 2012 Late Triassic bivalves and brachiopods from southern Mendoza, Argentina Revue de Paléobiologie, Genève spécial 11 317–344 http://www.ville-ge.ch/mhng/paleo/paleo-pdf/spec-11/pal-spec-11-22.pdf Sp. novValidManceñido in Damborenea & ManceñidoLate TriassicArgentinaA lepismatinid brachiopod, a species of Bolilaspirifer.Burrirhynchia angustisinuataT. N. Smirnova 2012 Early Cretaceous rhynchonellids of Dagestan: System, morphology, stratigraphic and paleobiogeographic significance Paleontological Journal 46 11 1197–1296 10.1134/S0031030112110019 2012PalJ...46.1197S 128422642 sp novValidSmirnovaEarly CretaceousRussiaA cyclothyridid rhynchonellide, a species of Burrirhynchia.Burrirhynchia latimarginatasp novValidSmirnovaEarly CretaceousRussiaA cyclothyridid rhynchonellide, a species of Burrirhynchia.Burrirhynchia owenisp novValidSmirnovaEarly CretaceousRussiaA cyclothyridid rhynchonellide, a species of Burrirhynchia.Burrirhynchia subgrasianasp novValidSmirnovaEarly CretaceousRussiaA cyclothyridid rhynchonellide, a species of Burrirhynchia.CallositellaGen. et sp. novValidRasmussen, Harper & BlodgettLate OrdovicianFarewell TerraneUnited StatesA member of Orthida belonging to the group Enteletoidea and the family Saukrodictyidae. The type species is C. cheeneetnukensis.Christiania aseptataSp. novValidRasmussen, Harper & BlodgettLate OrdovicianFarewell TerraneUnited StatesA member of Strophomenida belonging to the family Christianiidae.CincinnetinaJisuo Jin 2012 Cincinnetina, a new Late Ordovician dalmanellid brachiopod from the Cincinnati type area, USA: implications for the evolution and palaeogeography of the epicontinental fauna of Laurentia Palaeontology 55 1 205–228 10.1111/j.1475-4983.2011.01113.x 2012Palgy..55..205J 129264467 free Gen, sp. et comb. novValidJinLate OrdovicianUnited StatesA dalmanellid brachiopod, with C. multisecta, C. meeki, C. minnesotensisCraspedelia potterellaSp. novValidRasmussen, Harper & BlodgettLate OrdovicianFarewell TerraneUnited StatesA member of Strophomenida belonging to the group Plectambonitoidea and the family Bimuriidae.Crurithyris tazawaiWeihong He G. R. Shi Yang Zhang Tinglu Yang Fei Teng Shunbao Wu 2012 Systematics and palaeoecology of Changhsingian (Late Permian) Ambocoeliidae brachiopods from South China and implications for the end-Permian mass extinction Alcheringa: An Australasian Journal of Palaeontology 36 4 515–530 10.1080/03115518.2012.688669 2012Alch...36..515H 140157112 sp novValidHe et al..Late Permian (Changhsingian)ChinaAn ambocoeliid brachiopod, a species of Crurithyris.Cyclothyris aliformissp novValidSmirnovaEarly CretaceousRussiaA cyclothyridid rhynchonellide, a species of Burrirhynchia.Cyclothyris alikenticasp novValidSmirnovaEarly CretaceousRussiaA cyclothyridid rhynchonellide, a species of Burrirhynchia.Cyclothyris burgemakensissp novValidSmirnovaEarly CretaceousRussiaA cyclothyridid rhynchonellide, a species of Burrirhynchia.Cyclothyris dagestanicasp novValidSmirnovaEarly CretaceousRussiaA cyclothyridid rhynchonellide, a species of Burrirhynchia.Cyclothyris tenuicostatasp novValidSmirnovaEarly CretaceousRussiaA cyclothyridid rhynchonellide, a species of Burrirhynchia.Cyclothyris zudakharicasp novValidSmirnovaEarly CretaceousRussiaA cyclothyridid rhynchonellide, a species of Burrirhynchia.Daghanirhynchia susanaeSpeciesValidFeldman et al.CallovianMughanniyya FormationJordanA tetrarhynchiid rhynchonellide, a species of Daghanirhynchia.DuolobellaGen. et sp. novValidRasmussen, Harper & BlodgettLate OrdovicianFarewell TerraneUnited StatesA member of Orthida belonging to the group Orthoidea and the family Orthidae. The type species is D. sandiae.Eodmitria anaurisN. V. Oleneva 2012 New Spiriferids (Brachiopoda) from the Upper Devonian of Middle Timan Paleontological Journal 46 5 461–469 10.1134/S0031030112050073 2012PalJ...46..461O 128480437 sp novValidOlenevaLate Devonian (middle Frasnian)RussiaA cyrtospiriferid spiriferid, a species of Eodmitria.GatosellaJuan L. Benedetto 2012 Gatosella, a new basal plectambonitoid brachiopod with undercut cardinal process from Middle Ordovician limestones of the Precordillera terrane, Argentina Journal of Systematic Palaeontology 10 3 435–443 10.1080/14772019.2011.590536 2012JSPal..10..435B 128954396 11336/52205 free Gen. et sp. novValidBenedettoMiddle Ordovician (early Darriwilian)San Juan FormationArgentinaA plectambonitoid brachiopod. The type species is Gatosella muricata.Gelidorthis perisiberiaensisSp. novValidRasmussen, Harper & BlodgettLate OrdovicianFarewell TerraneUnited StatesA member of Orthida belonging to the group Plectorthoidea and the family Plectorthidae.Gracianella (Sublepida) paululaAndrzej Baliński 2012 The brachiopod succession through the Silurian–Devonian boundary beds at Dnistrove, Podolia, Ukraine Acta Palaeontologica Polonica 57 4 897–924 http://app.pan.pl/article/item/app20110138.html 10.4202/app.2011.0138 2012AcPaP..57..897B 55166314 free sp novValidBalińskiEarly DevonianKhudykivtsi BedsUkraineAn atrypinid brachiopod, a species of Gracianella.Kentronetes giaePatrick R. Racheboeuf Jean-Georges Casier Yves Plusquellec Margarita Toro David Mendoza Maria da Gloria Pires de Carvalho Alain Le Hérissé Florentin Paris Esperanza Fernández-Martinez Francis Tourneur Jean Broutin Sylvie Crasquin Philippe Janvier 2012 New data on the Silurian–Devonian palaeontology and biostratigraphy of Bolivia Bulletin of Geosciences 87 2 269–314 10.3140/bull.geosci.1248 free Sp. novValidRacheboeuf in Racheboeuf et al.Devonian (Lochkovian)Uncía FormationBoliviaA chonetoid brachiopod belonging to the family Strophochonetidae.KuangshanotretaHaizhou Wang Zhifei Zhang Lars E. Holmer Shixue Hu Xiangren Wang Guoxiang Li 2012 Peduncular attached secondary tiering acrotretoid brachiopods from the Chengjiang fauna: Implications for the ecological expansion of brachiopods during the Cambrian explosion Palaeogeography, Palaeoclimatology, Palaeoecology 323–325 60–67 10.1016/j.palaeo.2012.01.027 2012PPP...323...60W Gen. et sp. novValidWang et al.Early CambrianHeilinpu FormationChinaAn acrotretoid, a member of Lingulata. The type species is Kuangshanotreta malungensis.KudrjavzevinaGen. et 3 sp. novValidSmirnovaEarly CretaceousRussiaA cyclothyridid rhynchonellide. Genus contains 3 species: K. ulluchaensis, K. regularis and K. fragilis.Lamellaerhynchia latiovalisSp. novValidSmirnovaEarly CretaceousRussiaA praecyclothyridid rhynchonellide, a species of Lamellaerhynchia.Leptaena (Septomena) alaskensisSp. novValidRasmussen, Harper & BlodgettLate OrdovicianFarewell TerraneUnited StatesA member of Strophomenida belonging to the family Rafinesquinidae.LobachevinaGen. et 2 sp. novValidSmirnovaEarly CretaceousRussiaA cyclothyridid rhynchonellide. Genus contains 2 new species: L. angusteinis and L. vagus.LynnicaA. A. Madison 2012 Formation of cardinal process in Ordovician strophomenids Paleontological Journal 46 12 1362–1374 10.1134/S0031030112120039 2012PalJ...46.1362M 84981929 Gen. et sp. novValidMadisonOrdovicianRussiaA member of Strophomenida. The type species is Lynnica fragilis.MergliaChristian Franke 2012 Marine Fauna der Wiltz-Schichten (Ober-Emsium, Unter-Devon) der Mulde von Wiltz und der Daleider Mulden-Gruppe (Luxemburg, Deutschland). Teil 3: Craniida Ferrantia 68 133–147 https://ps.mnhn.lu/ferrantia/publications/Ferrantia68.pdf Gen. et comb. novValidFrankeDevonianGermany LuxembourgA member of the family Craniidae. The type species is "Philhedra" schwerdi Drevermann (1902).Orthothrix sudoiJun-ichi Tazawa 2012 Late Permian (Changhsingian) brachiopod fauna from Nabekoshiyama in the Kesennuma area, South Kitakami Belt, northeast Japan Science Reports of Niigata University. (Geology) 27 15–50 10191/22748 Sp. novValidTazawaPermian (Changhsingian)Toyoma FormationJapanA member of Productida belonging to the suborder Strophalosiidina and the family Strophalosiidae.Oanduporella kuskokwimensisSp. novValidRasmussen, Harper & BlodgettLate OrdovicianFarewell TerraneUnited StatesA member of Orthida belonging to the group Enteletoidea and the family Draboviidae.PalaeowingellaGen. et sp. novValidRasmussen, Harper & BlodgettLate OrdovicianFarewell TerraneUnited StatesA member of Orthida belonging to the group Orthoidea and the family Orthidae. The type species is P. farewellensis.Petrocrania fabisziskyiSp. novValidFrankeDevonianGermany LuxembourgA member of the family Craniidae.Petrocrania krautscheidensisSp. novValidFrankeDevonianGermany LuxembourgA member of the family Craniidae.Plectodonta mariae pantheraeSubsp novValidBalińskiEarly DevonianKhudykivtsi BedsUkraineA sowerbyellid brachiopod, a subspecies of Plectodonta mariae.Ptychoglyptus alaensisSp. novValidRasmussen, Harper & BlodgettLate OrdovicianFarewell TerraneUnited StatesA member of Strophomenida belonging to the group Plectambonitoidea and the family Sowerbyellidae.RotutaspiriferGen. et sp. novValidOlenevaLate Devonian (middle Frasnian)RussiaA cyrtospiriferid spiriferid. The type species is Rotutaspirifer rotutus.Sanjuanetes glemareciSp. novValidRacheboeuf in Racheboeuf et al.Silurian (Přídolí)Muruhuta Shale MemberBoliviaA chonetoid brachiopod belonging to the family Strophochonetidae.SharovaellaA. V. Pakhnevich 2012 New Devonian punctate rhynchonellids (Brachiopoda) from Transcaucasia Paleontological Journal 46 6 560–567 10.1134/S0031030112060081 2012PalJ...46..560P 129912631 Gen. et sp. novValidPakhnevichLate Devonian (Famennian)RussiaA punctate rhynchonellide brachiopod. The type species is Sharovaella mirabilis.Skenidioides tatyanaesp novValidBalińskiEarly DevonianKhudykivtsi BedsUkraineA skenidiid brachiopod, a species of Skenidioides.Sowerbyella (Rugosowerbyella) praecursorSp. novValidRasmussen, Harper & BlodgettLate OrdovicianFarewell TerraneUnited StatesA member of Strophomenida belonging to the group Plectambonitoidea and the family Sowerbyellidae.Sowerbyella (Sowerbyella) rectangularisSp. novValidRasmussen, Harper & BlodgettLate OrdovicianFarewell TerraneUnited StatesA member of Strophomenida belonging to the group Plectambonitoidea and the family Sowerbyellidae.StrophatrypaArthur J. Boucot Robert B. Blodgett David M. Rohr 2012 Strophatrypa, a new genus of Brachiopoda (Atrypidae), from upper Silurian strata of the Alexander terrane, northeast Chichagof Island, Alaska Bulletin of Geosciences 87 2 261–267 10.3140/bull.geosci.1285 free Gen. et sp. novValidBoucot, Blodgett & RohrLate SilurianUnited StatesA member of Atrypida belonging to the family Atrypidae. The type species is S. skaflestadi.Sulcirhynchia insolitusSp. novValidSmirnovaEarly CretaceousRussiaA praecyclothyridid rhynchonellide, a species of Sulcirhynchia.Terrakea nabekoshiyamensisSp. novValidTazawaPermian (Changhsingian)Toyoma FormationJapanA member of Productida belonging to the suborder Productidina and the family Linoproductidae.TransridgeiaGen. et sp. novValidRasmussen, Harper & BlodgettOrdovician (late Darriwilian–early Sandbian)Farewell TerraneUnited StatesA member of Strophomenida belonging to the family Strophomenidae. The type species is T. costata. Molluscs 2012 in molluscan paleontology The following is a summary of the Molluscs described in 2012 54 ammonites 14 other cephalopods 52 gastropods 15 other molluscs Echinoderms Name Novelty Status Authors Age Unit Location Notes ImagesAnatifopsis fillmorensisColin D. Sumrall James Sprinkle Thomas E. Guensburg Benjamin F. Dattilo 2012 Early Ordovician Mitrates and A Possible Solute (Echinodermata) from the Western United States Journal of Paleontology 86 4 595–604 http://jpaleontol.geoscienceworld.org/content/86/4/595.abstract 10.1666/10-165R.1 2012JPal...86..595S 131176256 subscription Sp. novValidSumrall et al.Early OrdovicianUnited StatesA mitrate, a species of Anatifopsis.Anatifopsis ninemilensisSp. novValidSumrall et al.Early OrdovicianMcKelligon Canyon Formation Ninemile ShaleUnited StatesA mitrate, a species of Anatifopsis.AndymetraHans Hess 2012 Crinoids from the Middle Jurassic (Bajocian–Lower Callovian) of Ardèche, France Swiss Journal of Palaeontology 131 2 211–253 10.1007/s13358-012-0044-9 2012SwJP..131..211H 129530026 Gen. et sp. novValidHessEarly BathonianFranceA comatulid crinoid. The type species is Andymetra galei.Aorocrinus meyeriWilliam I. Ausich Edgar W. Roeser 2012 Camerate and Disparid Crinoids from the Late Kinderhookian Meadville Shale, Cuyahoga Formation of Ohio Journal of Paleontology 86 3 488–507 http://jpaleontol.geoscienceworld.org/content/86/3/488.abstract 10.1666/11-102.1 2012JPal...86..488A 130268870 subscription Sp. novValidAusich & RoeserLate KinderhookianCuyahoga FormationUnited StatesA camerate crinoid, a species of Aorocrinus.Aphelecrinus gracilisSp. novValidKammer & RoeserKinderhookianCuyahoga FormationUnited StatesA cladid crinoid, a species of Aphelecrinus.Apiocrinites negevensisWilliam I. Ausich Mark A. Wilson 2012 New Tethyan Apiocrinitidae (Crinoidea, Articulata) from the Jurassic of Israel Journal of Paleontology 86 6 1051–1055 10.1666/12-049R.1 2012JPal...86.1051A 128734493 Sp. novValidAusich & WilsonMiddle Jurassic (Callovian)Matmor FormationIsraelAn apiocrinitid crinoid, a species of Apiocrinites.Aryballocrinus martiniSp. novValidAusich & RoeserLate KinderhookianCuyahoga FormationUnited StatesA camerate crinoid, a species of Aryballocrinus.Atelestocrinus meszarosiSp. novValidKammer & RoeserKinderhookianCuyahoga FormationUnited StatesA cladid crinoid, a species of Atelestocrinus.Brissopsis (Kleinia) riccardiiSara G. Parma 2012 El género Brissopsis L. Agassiz, 1840 (Echinoidea: Spatangoida) en el Paleógeno y Neógeno de la República Argentina Revue de Paléobiologie, Genève spécial 11 417–427 http://www.ville-ge.ch/mhng/paleo/paleo-pdf/spec-11/pal-spec-11-29.pdf Sp. novValidParmaEocene to Oligocene or early MioceneArgentinaA brissopsid echinoid, a species of Brissopsis.Calceocrinus balticensisWilliam I. Ausich Mark A. Wilson Olev Vinn 2012 Crinoids from the Silurian of Western Estonia Acta Palaeontologica Polonica 57 3 613–631 10.4202/app.2010.0094 2012AcPaP..57..613A 56571107 free Sp. novValidAusich, Wilson & VinnSilurian (Llandovery)EstoniaA disparid crinoid belonging to the group Calceocrinida.CtenoimbricataSamuel Zamora Imran A. Rahman Andrew B. Smith 2012 Plated Cambrian Bilaterians Reveal the Earliest Stages of Echinoderm Evolution PLOS ONE 7 6 e38296 10.1371/journal.pone.0038296 22701623 3368939 2012PLoSO...738296Z free Gen. et sp. novValidZamora, Rahman & SmithCambrianSpainA stem group echinoderm. The type species is Ctenoimbricata spinosa.CuyahogacrinusThomas W. Kammer Edgar W. Roeser 2012 Cladid Crinoids from the Late Kinderhookian Meadville Shale, Cuyahoga Formation of Ohio Journal of Paleontology 86 3 470–487 http://jpaleontol.geoscienceworld.org/content/86/3/470.abstract 10.1666/11-101.1 2012JPal...86..470K 140610304 subscription Gen. et sp. novValidKammer & RoeserKinderhookianCuyahoga FormationUnited StatesA cladid crinoid. The type species is Cuyahogacrinus lodiensis.Cyathocrinites simplexSp. novValidKammer & RoeserKinderhookianCuyahoga FormationUnited StatesA cladid crinoid, a species of Cyathocrinites.Cyrtocrinus praenutansSp. novValidHessEarly BathonianFranceA cyrtocrinid crinoid, a species of Cyrtocrinus.Desmidocrinus laevigatusSp. novValidAusich, Wilson & VinnSilurian (Pridoli)Kaugatuma FormationEstoniaA camerate crinoid belonging to the group Monobathrida.Diploblastus fadigaiJames W. Atwood Colin D. Sumrall 2012 Morphometric Investigation of the Pentremites Fauna from the Glen Dean Formation, Kentucky Journal of Paleontology 86 5 813–828 10.1666/12-003.1 2012JPal...86..813A 129854763 Sp. novValidAtwood & SumrallCarboniferous (Mississippian)Glen Dean FormationUnited StatesA blastoid, a species of Diploblastus.DrepanocystisGen. et sp. novValidSumrall et al.Early OrdovicianWah Wah LimestoneUSAA possible solute. The type species is Drepanocystis dubius.Eucalyptocrinites tumidusSp. novValidAusich, Wilson & VinnSilurian (Pridoli)Kaugatuma FormationEstoniaA camerate crinoid belonging to the group Monobathrida.Faorina maulluiP. Stara E. Borghi 2012 First fossil record of the genus Faorina (Echinoidea: Pericosmidae) Bollettino della Società Paleontologica Italiana 51 2 85–98 http://paleoitalia.org/archives/bollettino-spi/66/vol-51-2-2012/ 2013-01-28 2014-01-01 https://web.archive.org/web/20140101012306/http://paleoitalia.org/archives/bollettino-spi/66/vol-51-2-2012/ usurped Sp. novValidStara & BorghiEarly MioceneItalyA pericosmid sea urchin, a species of Faorina.Gastrocrinus leunisseniChristian Franke 2012 Marine Fauna der Wiltz-Schichten (Ober-Emsium, Unter-Devon) der Mulde von Wiltz und der Daleider Mulden-Gruppe (Luxemburg, Deutschland). Teil 2: Crinoida Ferrantia 68 5–64 https://ps.mnhn.lu/ferrantia/publications/Ferrantia68.pdf Sp. novValidFrankeDevonianGermanyA cladid crinoid belonging to the family Botryocrinidae.Goniocrinus sceletusSp. novValidKammer & RoeserKinderhookianCuyahoga FormationUnited StatesA cladid crinoid, a species of Goniocrinus.HeckerocrinusAlexander B. Doweld 2012 Heckerocrinus, a new substitute generic name for fossil crinoid Bockia Hecker 1940 (Echinodermata: Crinoidea) non Reisinger 1924 (Vermes: Turbellaria) Paläontologische Zeitschrift 86 4 457–458 10.1007/s12542-012-0148-y 2012PalZ...86..457D 89232853 Nom nov.ValidDoweldOrdovicianA replacement name for the crinoid genus Bockia Hecker 1940.LeadagmaraBen Thuy Yoshiaki Ishida Eiji Doi Andreas Kroh 2013 New ophiacanthid brittle stars (Echinodermata: Ophiuroidea) from the Upper Triassic of Japan: first insights into the origin and evolution of an extant deep-sea group Journal of Systematic Palaeontology 11 5 515–530 10.1080/14772019.2012.702691 4 July 2025 2013JSPal..11..515T https://www.tandfonline.com/doi/abs/10.1080/14772019.2012.702691Gen. et sp. novValidhttp://zoobank.org/NomenclaturalActs/A079A3B1-1CA4-4C69-98D2-A89A4D8BE8C7 ZooBank.org zoobank.org 23 April 2018Thuy et al.Late Triassic (early Carnian)JapanAn ophiacanthid brittle star. The type species is Leadagmara gracilispina.Lebetocrinus ohioensisSp. novValidKammer & RoeserKinderhookianCuyahoga FormationUnited StatesA cladid crinoid, a species of Lebetocrinus.NeotaxocrinusG. V. Mirantsev 2012 A new taxocrinid genus (Crinoidea, Flexibilia) from the Upper Carboniferous of the Moscow Region Paleontological Journal 46 6 568–574 10.1134/S0031030112060068 2012PalJ...46..568M 131544886 Gen. et sp. novValidMirantsevLate Carboniferous (Pennsylvanian)RussiaA taxocrinid crinoid. The type species is Neotaxocrinus arendti.Palaeocomaster messingiSp. novValidHessEarly BathonianFranceA comatulid crinoid, a species of Palaeocomaster.Paradiabolocrinus teresPaul W. Hearn Bradley Deline 2012 A new species of Paradiabolocrinus from the Upper Ordovician of central Kentucky, USA Southeastern Geology 49 1 25–30 https://www.academia.edu/24986865 Sp. novValidHearn & DelineLate OrdovicianLexington LimestoneUnited StatesA camerate crinoid belonging to the group Diplobathra and the family Rhodocrinitidae.Pentacrinites ausichiSp. novValidHessLate Bajocian or early BathonianFranceA species of Pentacrinites.Pentremites frederickiSp. novValidAtwood & SumrallCarboniferous (Mississippian)Glen Dean FormationUnited StatesA blastoid, a species of Pentremites.Pentremites meganaeSp. novValidAtwood & SumrallCarboniferous (Mississippian)Glen Dean FormationUnited StatesA blastoid, a species of Pentremites.Phyllocrinus voultensisSp. novValidHessEarly BathonianFranceA cyrtocrinid crinoid, a species of Phyllocrinus.Phymosoma ravniNils Schlüter Manfred Kutscher Andrew B. Smith John W. M. Jagt Jackie A. Lees 2012 Late Cretaceous phymosomatids and the true identity of Cidarites granulosus Goldfuss, 1829 (Echinoidea, Phymosomatoida) Zootaxa 3271 17–30 http://www.mapress.com/zootaxa/2012/f/z03271p030f.pdf 10.11646/zootaxa.3271.1.2 Sp nov.ValidSchlüter et al.Late CretaceousWestern Europe and Middle EastA phymosomatoid sea urchin, a species of Phymosoma.Platycrinites burkeiSp. novValidAusich & RoeserLate KinderhookianCuyahoga FormationUnited StatesA camerate crinoid, a species of Platycrinites sensu lato.Praetetracrinus bathonicusSp. novValidHessEarly BathonianFranceA cyrtocrinid crinoid, a species of Praetetracrinus.Propoteriocrinus hosingeniSp. novValidFrankeDevonianGermany LuxembourgA cladid crinoid belonging to the family Glossocrinidae.PygolampasThomas Saucède Jean-Christophe Dudicourt Philippe Courville 2012 Description of two new fossil echinoids (Echinodermata: Echinoidea) from the Early Hauterivian (Early Cretaceous) of the Paris Basin (France) Zootaxa 3512 75–88 http://www.mapress.com/zootaxa/2012/f/z03512p088f.pdf 10.11646/zootaxa.3512.1.5 Gen. et sp. novValidSaucède, Dudicourt & CourvilleEarly Cretaceous (Early Hauterivian)FranceA sea urchin. The type species is Pygolampas edita.SaaremaacrinusGen. et sp. novValidAusich, Wilson & VinnSilurian (Pridoli)Kaugatuma FormationEstoniaA camerate crinoid belonging to the group Monobathrida. The type species is S. estoniensis.SalvasterGen. et sp. novValidSaucède, Dudicourt & CourvilleEarly Cretaceous (Early Hauterivian)FranceA sea urchin. The type species is Salvaster roberti.ScutellacrinusGen. et sp. novValidHessLate Bajocian or early BathonianFranceA cyrtocrinid crinoid. The type species is Scutellacrinus tenuis.Segmentocolumnus (col.) hanshessiStephen K. Donovan David A. T. Harper Zhan Renbin Lars Stemmerik Liu Jianbo Svend Stouge 2012 A primitive cladid crinoid from the Jiacun Group, Tibet (Darriwilian, Middle Ordovician) Geological Journal 47 6 653–660 10.1002/gj.2436 2012GeolJ..47..653D 129177823 https://durham-repository.worktribe.com/output/1506057 Sp. novValidDonovan et al.Middle Ordovician (latest Darriwilian)ChinaProbably a stem-group cladid crinoid, possibly a dendrocrinid; a species of Segmentocolumnus.SingillatimetraGen. et sp. novValidHessEarly BathonianFranceA comatulid crinoid. The type species is Singillatimetra inordinata.Solanocrinites voultensisSp. novValidHessEarly BathonianFranceA comatulid crinoid, a species of Solanocrinites.Thylechinus (Thylechinus) sinaiensisMarouf Abdel-Aty Mohamed Abdelhamid Mahmoud Moustafa Azab 2012 Turonian-Santonian echinoids from Egypt Geodiversitas 34 3 575–615 http://www.mnhn.fr/museum/front/medias/publication/46737_g2012n3a7.pdf 10.5252/g2012n3a7 2012Geodv..34..575M 198151884 April 2019 InternetArchiveBot yes Sp. novValidAbdelhamid & AzabTuronianEgyptA phymosomatid echinoid, a species of Thylechinus.Toxaster dakhlensisSp. novValidAbdelhamid & AzabTuronianEgyptA toxasterid echinoid, a species of Thylechinus.TripatagusLouis G. Zachos 2012 A new genus and species of echinoid (Echinoidea, Spatangoida) from the Oligocene (Rupelian) of Mississippi Southeastern Geology 49 1 43–48 https://www.researchgate.net/publication/236737020 Gen. et sp. novValidZachosOligocene (Rupelian)Marianna LimestoneUnited StatesA heart urchin belonging to the group Micrasterina. The type species is T. pittsi.YanjiahellaJunfeng Guo Yong Li Huiping Han Qiang Ou Jianren Zhou Yajuan Zheng 2012 New Macroscopic Problematic Fossil from the Early Cambrian Yanjiahe Biota, Yichang, Hubei, China Acta Geologica Sinica (English Edition) 86 4 791–798 http://www.geojournals.cn/dzxben/ch/reader/view_abstract.aspx?file_no=201204001&flag=1 10.1111/j.1755-6724.2012.00706.x 2012AcGlS..86..791G 84790682 subscription Gen. et 3 sp. novValidGuo et al.Early CambrianYanjiahe FormationChinaAn animal of uncertain phylogenetic position, a possible relative of echinoderms. Genus contains Yanjiahella ancarpa (the type species), Y. monocarpa and Y. biscarpa. Ascidians Name Novelty Status Authors Age Unit Location Notes ImagesBurykhiaM. A. Fedonkin P. Vickers-Rich B. J. Swalla P. Trusler M. Hall 2012 A new metazoan from the Vendian of the White Sea, Russia, with possible affinities to the ascidians Paleontological Journal 46 1 1–11 10.1134/S0031030112010042 2012PalJ...46....1F 128415270 Gen. et sp.ValidFedonkin et al.VendianVerkhovka Formation or Ust-Pinega FormationRussiaAn ausiid, a possible ascidian, Its type species is Burykhia hunti. Conodonts Name Novelty Status Authors Age Unit Location Notes ImagesCarnepigondolella angulataMichele Mazza Andrea Cau Manuel Rigo 2012 Application of numerical cladistic analyses to the Carnian–Norian conodonts: a new approach for phylogenetic interpretations Journal of Systematic Palaeontology 10 3 401–422 10.1080/14772019.2011.573584 2012JSPal..10..401M 2434/223494 53501178 free Sp novValidMazza, Cau & RigoLate TriassicCanada ItalyOriginally described as a species of Carnepigondolella; Orchard (2014) transferred this species to the genus Quadralella.Michael J. Orchard 2014 Conodonts from the Carnian-Norian Boundary (Upper Triassic) of Black Bear Ridge, Northeastern British Columbia, Canada New Mexico Museum of Natural History and Science Bulletin 64 1–139 Carnepigondolella gulloaeMichele Mazza Manuel Rigo Maria Gullo 2012 Taxonomy and biostratigraphic record of the Upper Triassic conodonts of the Pizzo Mondello section (western Sicily, Italy), GSSP candidate for the base of the Norian Rivista Italiana di Paleontologia e Stratigrafia 118 1 85–130 http://www.rivistaitalianadipaleontologia.it/pub/index.php/abstracts-vol-118-1-2012 2015-04-01 https://web.archive.org/web/20150402175107/http://www.rivistaitalianadipaleontologia.it/pub/index.php/abstracts-vol-118-1-2012 2015-04-02 Sp novValidMazza, Rigo & GulloLate TriassicScillato FormationItalyA species of Carnepigondolella.Carnepigondolella tuvalicaSp novValidMazza, Rigo & GulloLate TriassicScillato FormationItalyA species of Carnepigondolella.Caudicriodus schoenlaubiDaniel Drygant Hubert Szaniawski 2012 Lochkovian conodonts from Podolia, Ukraine, and their stratigraphic significance Acta Palaeontologica Polonica 57 4 833–861 http://app.pan.pl/article/item/app20120124.html 10.4202/app.2012.0124 free Sp novValidDrygant & SzaniawskiEarly Devonian (middle to late Lochkovian)Chortkiv FormationUkraineAn icriodontid, a species of Caudicriodus.Epigondolella heinziSp novValidMazza, Cau & RigoLate TriassicItalyA species of Epigondolella.Epigondolella miettoiSp novValidMazza, Cau & RigoLate TriassicItalyA species of Epigondolella.Idiognathodus espinamaenisC. A. Méndez 2012 Moscovian conodonts from the Nevandi River Valley (Cantabrian Zone, North Spain) Trabajos de Geología 32 63–75 http://www.unioviedo.es/reunido/index.php/TDG/article/view/9886 Sp. novValidMéndezCarboniferous (Moscovian)SpainIdiognathodus mendeziSp. novValidMéndezCarboniferous (Moscovian)SpainMeiognathusShu-zhong Shen Dong-xun Yuan Charles M. Henderson Junichi Tazawa Yi-chun Zhang 2012 Implications of Kungurian (Early Permian) conodonts from Hatahoko, Japan, for correlation between the Tethyan and international timescales Micropaleontology 58 6 505–522 10.47894/mpal.58.6.03 24413308 2012MiPal..58..505S 127120834 Gen. et sp. novValidShen et al.Permian (Kungurian)JapanA member of Ozarkodinida belonging to the family Sweetognathidae. The type species is Meiognathus pustulus.MuellerinaI. A. Bardashev N. P. Bardasheva 2012 Platformennye konodonty iz pogranichnykh zhivet-franskikh otlozhenii (srednii-verkhnii devon) Donish 1–90 Gen. et sp. novJunior homonymBardashev & BardashevaMiddle Devonian (Givetian)TajikistanThe type species is Muellerina idrisovi. The generic name turned out to be preoccupied by Muellerina Bassiouni (1965); Bardashev & Bardasheva (2013) subsequently renamed the conodont genus Muellerilepis.I. A. Bardashev N. P. Bardasheva 2013 Muellerilepis, a new replacement generic name for Muellerina Bardashev et Bardasheva, 2012 (Conodonta) Paleontological Journal 47 5 554 10.1134/S0031030113050055 2013PalJ...47..554B 84383505 free NealeodusGen. et comb. novValidStougeMiddle OrdovicianCanadaA new genus for "Lenodus" martinpointensis (Johnston and Barnes, 2000).Norigondolella trinacriaeSp novValidMazza, Cau & RigoLate TriassicItalyOriginally described as a species of Norigondolella; Karádi, Kozur & Görögt (2013) considered it more likely to be a species of Paragondolella.Viktor Karádi Heinz W. Kozur Ágnes Görögt 2013 Stratigraphically important lower Norian conodonts from the Csővár borehole (CSV-1), Hungary – Comparison with the conodont succession of the Norian GSSP candidate Pizzo Mondello New Mexico Museum of Natural History and Science Bulletin 61 284–295 http://paleo.cortland.edu/globaltriassic2/Bulletin%2061%20Final/21-Karadi%20et%20al%20(Pizzomondello).pdf 2017-11-02 2017-11-07 https://web.archive.org/web/20171107003009/http://paleo.cortland.edu/globaltriassic2/Bulletin%2061%20Final/21-Karadi%20et%20al%20(Pizzomondello).pdf Pandorinellina? parvaSp novValidDrygant & SzaniawskiEarly Devonian (late Lochkovian or Pragian)Ivanye FormationUkraineA spathognathodontid, possibly a species of Pandorinellina.Periodon hankensisSvend Stouge 2012 Middle Ordovician (late Dapingian–Darriwilian) conodonts from the Cow Head Group and Lower Head Formation, western Newfoundland, Canada Canadian Journal of Earth Sciences 49 1 59–90 10.1139/e11-057 2012CaJES..49...59S Sp novValidStougeMiddle OrdovicianCanadaA species of Periodon.Polygnathus chongqingensisLiming Gong Chengyuan Wang Changsheng Wang Yunxi Hu Yongyang Tang Ping Hu Qingdong Wang 2012 Devonian conodonts in SE area of Chongqing City and its geological significance Acta Micropalaeontologica Sinica 29 3 282–298 http://eng.oversea.cnki.net/kcms/detail/detail.aspx?filename=WSGT201203008&DBName=cjfqtotal&dbcode=cjfq 2016-02-06 2016-02-06 https://web.archive.org/web/20160206232345/http://eng.oversea.cnki.net/kcms/detail/detail.aspx?filename=WSGT201203008&DBName=cjfqtotal&dbcode=cjfq Sp. novValidWang in Gong et al.DevonianChinaPseudopolygnathus inordinatusHarald Tragelehn Sven Hartenfels 2012 Pseudopolygnathus inordinatus nov. nom. – homonym replacement for an upper Famennian (Upper Devonian) conodont species Subcommission on Devonian Stratigraphy Newsletter 27 59–60 http://unica2.unica.it/sds/images/SDS27%20-%202012.pdf Nom. novValidTragelehn & HartenfelsDevonian (Famennian)GermanyA replacement name for Pseudopolygnathus irregularis Tragelehn & Hartenfels (2011) (preoccupied by Pseudopolygnathus irregularis Branson, 1934).Spinodus wardiSp novValidStougeMiddle OrdovicianCanadaA species of Spinodus. Fishes 2012 in fish paleontology During 2012, 81 new species of fish were described. Amphibians Research A study of the braincase of Eocaecilia micropodia and a phylogenetic analysis of non-amniote tetrapods is published by Hillary C. Maddin, Farish A. Jenkins Jr and Jason S. Anderson (2012).Hillary C. Maddin Farish A. Jenkins Jr Jason S. Anderson 2012 The Braincase of Eocaecilia micropodia (Lissamphibia, Gymnophiona) and the Origin of Caecilians PLOS ONE 7 12 e50743 10.1371/journal.pone.0050743 23227204 3515621 2012PLoSO...750743M free A study of anatomy and relationships of Solenodonsaurus janenschi is published by Marylène Danto, Florian Witzmann and Johannes Müller (2012).Marylène Danto Florian Witzmann Johannes Müller 2012 Redescription and phylogenetic relationships of Solenodonsaurus janenschi Broili, 1924, from the Late Carboniferous of Nýřany, Czech Republic Fossil Record 15 2 45–59 10.1002/mmng.201200003 free A study of limb joint mobility of Ichthyostega is published by Stephanie E. Pierce, Jennifer A. Clack and John R. Hutchinson (2012).Stephanie E. Pierce Jennifer A. Clack John R. Hutchinson 2012 Three-dimensional limb joint mobility in the early tetrapod Ichthyostega Nature 486 7404 523–526 10.1038/nature11124 22722854 2012Natur.486..523P 3127857 https://rvc-repository.worktribe.com/preview/1642603/The%20Laryngoscope%20-%202023%20-%20Kirsch.pdf New taxa Newly named basal tetrapods Name Novelty Status Authors Age Unit Location Notes ImagesYmeriaClack J.A. Ahlberg, P.E. Blom, H. Finney, S.M. 2012 A new genus of Devonian tetrapod from North-East Greenland, with new information on the lower jaw of Ichthyostega Palaeontology 55 1 73–86 10.1111/j.1475-4983.2011.01117.x 2012Palgy..55...73C free Gen. et sp. novValidClack, Ahlberg, Blom & FinneyFamennianCelsius Bjerg GroupGreenlandA stem tetrapod closely related to Ichthyostega. Newly named temnospondyls Name Novelty Status Authors Age Unit Location Notes ImagesArachanaGraciela Piñeiro Alejandro Ramos Claudia Marsicano 2012 A rhinesuchid-like temnospondyl from the Permo-Triassic of Uruguay Comptes Rendus Palevol 11 1 65–78 10.1016/j.crpv.2011.07.007 2012CRPal..11...65P Gen. et sp. novValidPiñeiro, Ramos & Marsicano? Late PermianBuena Vista FormationUruguayA rhinesuchid-like stereospondyl.Benthosuchus gusevaeI. V. Novikov 2012 New data on trematosauroid labyrinthodonts of Eastern Europe: 4. Genus Benthosuchus Efremov, 1937 Paleontological Journal 46 4 400–411 10.1134/S0031030112040089 2012PalJ...46..400N 84240080 Sp. novValidNovikovEarly TriassicKamennyi Yar FormationRussiaA trematosauroid.Cacops woehriFröbisch N.B. and Reisz, R.R. 2012 A new species of dissorophid (Cacops woehri) from the Lower Permian Dolese Quarry, near Richards Spur, Oklahoma Journal of Vertebrate Paleontology 32 1 35–44 10.1080/02724634.2012.633586 2012JVPal..32...35F 131513990 Sp. novValidFröbisch & ReiszLeonardianUSAA dissorophid.Microposaurus averyiAnne Warren 2012 The South African stereospondyl Microposaurus from the Middle Triassic of the Sydney Basin, Australia Journal of Vertebrate Paleontology 32 3 538–544 10.1080/02724634.2012.658934 2012JVPal..32..538W 128842227 Sp novValidWarrenAnisianRouse Hill SiltstoneAustraliaA trematosaurine stereospondyl, a species of Microposaurus.thumb|150px|center|Microposaurus averyi.NyranerpetonRalf Werneburg 2012 Dissorophoide Amphibien aus dem Westphalian D (Ober-Karbon) von Nýřany in Böhmen (Tschechische Republik) - der Schlüssel zum Verständnis der frühen 'Branchiosaurier' Semana. Naturwissenschaftliche Veröffentlichungen des Naturhistorischen Museums Schloss Bertholdsburg Schleusingen 27 3–50 https://www.researchgate.net/publication/270877129 Gen. et sp. novValidWerneburgLate CarboniferousCzech RepublicA micromelerpetontid.Rainer R. Schoch Andrew R. Milner Handbook of Paleoherpetology. Part 3A2. Temnospondyli I Verlag Dr. Friedrich Pfeil 45 978-3-89937-170-3 2014 The type species is Nyranerpeton amilneri.Platyrhinops fritschiSp. novValidWerneburgLate CarboniferousCzech RepublicAn amphibamiform. Originally described as a species of Platyrhinops, but subsequently made the type species of a separate genus Anthracobamus.Ralf Werneburg 2019 Dissorophoid amphibians from the Carboniferous-Permian boundary of France Semana. Naturwissenschaftliche Veröffentlichungen des Naturhistorischen Museums Schloss Bertholdsburg Schleusingen 34 11–51 https://www.researchgate.net/publication/337870091 QantasI. V. Novikov 2012 New data on trematosauroid labyrinthodonts of Eastern Europe: 3. Qantas samarensis gen. et sp. nov. Paleontological Journal 46 2 177–186 10.1134/S0031030112020098 2012PalJ...46..177N 85092152 Gen. et sp. novValidNovikovEarly TriassicKamennyi Yar FormationRussiaA trematosauroid temnospondyl. The type species is Qantas samarensis.Sclerocephalus stambergiJozef Klembara J. Sébastien Steyer 2012 A new species of Sclerocephalus (Temnospondyli: Stereospondylomorpha) from the Early Permian of the Boskovice Basin (Czech Republic) Journal of Paleontology 86 2 302–310 http://jpaleontol.geoscienceworld.org/content/86/2/302.abstract 10.1666/11-051.1 2012JPal...86..302K 130383748 subscription Sp novValidKlembara & SteyerEarly PermianBoskovice BasinCzech RepublicA stereospondylomorph, a species of Sclerocephalus.TrematosuchoidesGen. et sp. novValidNovikovTriassicCynognathus Assemblage ZoneSouth AfricaA trematosaurid temnospondyl. The type species is Trematosuchoides africanus. Newly named lepospondyls Name Novelty Status Authors Age Unit Location Notes ImagesAltenglanerpetonGlienke S. 2012 A new "microsaur" (Amphibia; Lepospondyli) from the Rotliegend of the Saar–Palatinate region (Carboniferous/Permian transition; West Germany) Paläontologische Zeitschrift 86 3 297–311 10.1007/s12542-012-0130-8 2012PalZ...86..297G 140699792 Gen. et sp. novValidGlienkeGzhelian or AsselianAltenglan FormationGermanyA microsaur with an elongated body. Newly named lissamphibians Name Novelty Status Authors Age Unit Location Notes ImagesAlbionbatrachus oligocenicusMárton Venczel Vlad Codrea Cristina Fărcaş 2013 A new palaeobatrachid frog from the early Oligocene of Suceag, Romania Journal of Systematic Palaeontology 11 2 179–189 10.1080/14772019.2012.671790 2013JSPal..11..179V 129158241 Sp. novValidhttp://zoobank.org/NomenclaturalActs/D582DF3D-14FA-48FE-8240-8B81727B358E ZooBank.org zoobank.org 23 April 2018Venczel, Codrea & FărcaşEarly OligoceneRomaniaA palaeobatrachid frog, a species of Albionbatrachus.BakonybatrachusZoltán Szentesi Márton Venczel 2012 A new discoglossid frog from the Upper Cretaceous (Santonian) of Hungary Cretaceous Research 34 327–333 10.1016/j.cretres.2011.11.012 2012CrRes..34..327S Gen. et sp. novValidSzentesi & VenczelSantonianCsehbánya FormationHungaryA discoglossine discoglossid.BeiyanerpetonKe-Qin Gao Neil H. Shubin 2012 Late Jurassic salamandroid from western Liaoning, China Proceedings of the National Academy of Sciences of the United States of America 109 15 5767–5772 10.1073/pnas.1009828109 22411790 3326464 2012PNAS..109.5767G free Gen. et sp. novValidGao & ShubinOxfordianTiaojishan FormationChinaA salamandroid. The type species is Beiyanerpeton jianpingensis.Calyptocephalella satanFederico Agnolin 2012 A new Calyptocephalellidae (Anura, Neobatrachia) from the Upper Cretaceous of Patagonia, Argentina, with comments on its systematic position http://revistas.usal.es/index.php/0211-8327/article/view/12820 Studia Geologica Salmanticensia 48 2 129–178 Sp. novValidAgnolinLate Cretaceous (Campanian-Maastrichtian)Allen FormationArgentinaA member of Calyptocephalellidae, a relative of the helmeted water toad.Gigantobatrachus casamiquelaiSp. novValidAgnolinPaleoceneArgentinaA member of Calyptocephalellidae, a species of Gigantobatrachus.Piceoerpeton nayloriJames D. Gardner 2012 Revision of Piceoerpeton Meszoely (Caudata: Scapherpetontidae) and description of a new species from the late Maastrichtian and ?early Paleocene of western North America Bulletin de la Société Géologique de France 183 6 611–620 10.2113/gssgfbull.183.6.611Sp. nov.ValidGardnerLatest Cretaceous (late Maastrichtian), possibly also early PaleoceneUnited StatesA scapherpetontid salamander, a species of Piceoerpeton.Rana auscitanaC. Martín M. A. Alonso-Zarazaga B. Sanchiz 2012 Nomenclatural notes on living and fossil amphibians Graellsia 68 1 159–180 10.3989/graellsia.2012.v68.056 free Nom. novValidMartín, Alonso-Zarazaga & SanchizMiddle MioceneFranceA replacement name for Rana pygmaea Lartet (1851).Rana cadurcorumNom. novJunior synonymMartín, Alonso-Zarazaga & SanchizEoceneQuercy PhosphoritesFranceA replacement name for Rana plicata Filhol (1877). Reinterpreted as a junior synonym of Thaumastosaurus gezei by Laloy et al. (2013).Fabien Laloy Jean-Claude Rage Susan E. Evans Renaud Boistel Nicolas Lenoir Michel Laurin 2013 A re-interpretation of the Eocene anuran Thaumastosaurus based on microCT examination of a 'mummified' specimen PLOS ONE 8 9 e74874 10.1371/journal.pone.0074874 24086389 3783478 2013PLoSO...874874L free Rana sendoaNom. novValidMartín, Alonso-Zarazaga & SanchizLate PleistoceneGermanyA replacement name for Rana robusta Brunner (1956).Seminobatrachus2012 A new salamander from the late Paleocene–early Eocene of Ukraine Acta Palaeontologica Polonica 57 1 135–148 10.4202/app.2010.0101 Pavel P. Skutschas Yuri M. Gubin 2012AcPaP..57..135S 55252741 free Gen. et sp. novValidSkutschas & Gubinlate Paleocene - early EoceneUkraineA urodelan salamanderUberabatrachus2012 The diverse Cretaceous neobatrachian fauna of South America: Uberabatrachus carvalhoi, a new frog from the Maastrichtian Marília Formation, Minas Gerais, Brazil Gondwana Research 22 3–4 1141–1150 10.1016/j.gr.2012.02.021 Ana M. Báez Raúl O. Gómez Luiz C.B. Ribeiro Agustín G. Martinelli Vicente P.A. Teixeira Mara L.F. Ferraz 2012GondR..22.1141B Gen. et sp. novValidBáez et al.MaastrichtianMarília FormationBrazilA neobatrachian, a possible member of Nobleobatrachia. The type species is Uberabatrachus carvalhoi. Parareptiles Newly named parareptiles Name Novelty Status Authors Age Unit Location Notes ImagesFeeserpetonMark J. Macdougall Robert Reisz 2012 A new parareptile (Parareptilia, Lanthanosuchoidea) from the Early Permian of Oklahoma Journal of Vertebrate Paleontology 32 5 1018–1026 10.1080/02724634.2012.679757 2012JVPal..32.1018M 86218618 Gen. et sp. nov.ValidMacdougall & ReiszEarly PermianUnited StatesA lanthanosuchoid. The type species is Feeserpeton oklahomensis.LasasaurusJocelyn Falconnet Misalalanirina Andriamihaja Émilie Läng J.-Sébastien Steyer 2012 First procolophonid (Reptilia, Parareptilia) from the Lower Triassic of Madagascar Comptes Rendus Palevol 11 5 357–369 10.1016/j.crpv.2012.04.001 2012CRPal..11..357F Gen. et sp. nov.ValidFalconnet et al.Lower TriassicMiddle Sakamena FormationMadagascarA procolophonid. The type species is Lasasaurus beltanae. Ichthyopterygians Newly named ichthyosaurs Name Novelty Status Authors Age Unit Location Notes ImagesAcamptonectesFischer, V. Maisch, M.W. Naish, D. Kosma, R. Liston, J. Joger, U. Krüger, F.J. Pérez, J.P. Tainsh, J. Appleby, R.M. 3 2012 New Ophthalmosaurid Ichthyosaurs from the European Lower Cretaceous Demonstrate Extensive Ichthyosaur Survival across the Jurassic–Cretaceous Boundary PLOS ONE 7 1 e29234 10.1371/journal.pone.0029234 22235274 3250416 2012PLoSO...729234F free Gen. et sp. novValidFischer et al..Hauterivian–CenomanianCambridge Greensand FormationUnited KingdomAn ophthalmosaurid. Type species is Acamptonectes densus.thumb|150px|center|Skull roof of Acamptonectes densus.CryopterygiusPatrick S. Druckenmiller Jørn H. Hurum Espen M. Knutsen Hans Arne Nakrem 2012 Two new ophthalmosaurids (Reptilia: Ichthyosauria) from the Agardhfjellet Formation (Upper Jurassic: Volgian/Tithonian), Svalbard, Norway Norwegian Journal of Geology 92 2–3 311–339 http://www.geologi.no/norwegian-journal-of-geology-91/details/1/61-61 Gen. et sp. novDisputedDruckenmiller et al.Late JurassicAgardhfjellet FormationNorwayAn ophthalmosaurid. The type species is Cryopterygius kristiansenae. Zverkov & Efimov (2019) considered this species to be synonymous with Undorosaurus gorodischensis,Nikolay G. Zverkov Vladimir M. Efimov 2019 Revision of Undorosaurus, a mysterious Late Jurassic ichthyosaur of the Boreal Realm Journal of Systematic Palaeontology 17 14 963–993 10.1080/14772019.2018.1515793 2019JSPal..17.1183Z 91912834 https://figshare.com/articles/Revision_of_i_Undorosaurus_i_a_mysterious_Late_Jurassic_ichthyosaur_of_the_Boreal_Realm/7624868 while Delsett et al. (2019) considered it to be a distinct species, albeit possibly belonging to the genus Undorosaurus.Lene L. Delsett Aubrey J. Roberts Patrick S. Druckenmiller Jørn H. Hurum 2019 Osteology and phylogeny of Late Jurassic ichthyosaurs from the Slottsmøya Member Lagerstätte (Spitsbergen, Svalbard) Acta Palaeontologica Polonica 64 4 717–743 10.4202/app.00571.2018 free 10852/76089 free PalvenniaGen. et sp. novValidDruckenmiller et al.Late JurassicAgardhfjellet FormationNorwayAn ophthalmosaurid. The type species is Palvennia hoybergeti. Zverkov & Prilepskaya (2019) considered Palvennia to be a junior synonym of the genus Arthropterygius, though the authors maintained P. hoybergeti as a distinct species within the latter genus;Nikolay G. Zverkov Natalya E. Prilepskaya 2019 A prevalence of Arthropterygius (Ichthyosauria: Ophthalmosauridae) in the Late Jurassic—earliest Cretaceous of the Boreal Realm PeerJ 7 e6799 10.7717/peerj.6799 31106052 6497043 free Delsett et al. (2019) rejected this synonymy.Stenopterygius aaleniensisErin E. Maxwell Marta S. Fernández Rainer R. Schoch 2012 First Diagnostic Marine Reptile Remains from the Aalenian (Middle Jurassic): A New Ichthyosaur from Southwestern Germany PLOS ONE 7 8 e41692 10.1371/journal.pone.0041692 22870244 3411580 2012PLoSO...741692M free Sp. novValidMaxwell, Fernández & SchochAalenianGermanyA species of Stenopterygius.thumb|150px|center|The holotype of Stenopterygius aaleniensis.Temnodontosaurus azerguensisJeremy E. Martin Valentin Fischer Peggy Vincent Guillaume Suan 2012 A longirostrine Temnodontosaurus (Ichthyosauria) with comments on Early Jurassic ichthyosaur niche partitioning and disparity Palaeontology 55 5 995–1005 10.1111/j.1475-4983.2012.01159.x 2012Palgy..55..995M free Sp. novValidMartin et al.Middle ToarcianFranceA species of Temnodontosaurus. Lepidosauromorphs Newly named saurosphargids Name Novelty Status Authors Age Unit Location Notes ImagesLargocephalosaurusLong Cheng Xiaohong Chen Xiongwei Zeng Yongjian Cai 2012 A new eosauropterygian (Diapsida: Sauropterygia) from the Middle Triassic of Luoping, Yunnan Province Journal of Earth Science 23 1 33–40 10.1007/s12583-012-0231-z 2012JEaSc..23...33C 131061655 Gen. et sp. novValidCheng, Chen, Zeng & CaiMiddle TriassicGuanling FormationChinaInitially thought to be an eosauropterygian sauropterygian and a relative of Wumengosaurus, pachypleurosaurs and nothosauroids, but subsequently reinterpreted as member of Saurosphargidae (a non-sauropterygian diapsid, though related to sauropterygians), closely related to Saurosphargis and Sinosaurosphargis.Chun Li Da-Yong Jiang Long Cheng Xiao-Chun Wu Olivier Rieppel 84534618 2014 A new species of Largocephalosaurus (Diapsida: Saurosphargidae), with implications for the morphological diversity and phylogeny of the group Geological Magazine 151 1 100–120 10.1017/S001675681300023X 2014GeoM..151..100L The type species is Largocephalosaurus polycarpon. Newly named sauropterygians Name Novelty Status Authors Age Unit Location Notes ImagesAlbertonectesTai Kubo Mark T. Mitchell Donald M. Henderson 2012 Albertonectes vanderveldei, a new elasmosaur (Reptilia, Sauropterygia) from the Upper Cretaceous of Alberta Journal of Vertebrate Paleontology 32 3 557–572 10.1080/02724634.2012.658124 2012JVPal..32..557K 129500470 Gen. et sp. novValidKubo, Mitchell & HendersonUpper CampanianBearpaw FormationCanadaAn elasmosaurid. The type species is Albertonectes vanderveldei.thumb|150px|center|Albertonectes vanderveldei.AnningasauraPeggy Vincent Roger B. J. Benson 2012 Anningasaura, a basal plesiosaurian (Reptilia, Plesiosauria) from the Lower Jurassic of Lyme Regis, United Kingdom Journal of Vertebrate Paleontology 32 5 1049–1063 10.1080/02724634.2012.686467 2012JVPal..32.1049V 86547069 Gen. et sp. nov.ValidVincent & BensonEarly JurassicUnited KingdomA basal member of Plesiosauria. The type species is Anningasaura lymense.AvalonnectesRoger B. J. Benson Mark Evans Patrick S. Druckenmiller 2012 High Diversity, Low Disparity and Small Body Size in Plesiosaurs (Reptilia, Sauropterygia) from the Triassic–Jurassic Boundary PLOS ONE 7 3 e31838 10.1371/journal.pone.0031838 22438869 3306369 2012PLoSO...731838B free Gen. et sp. novValidBenson, Evans & DruckenmillerEarly Jurassic, most likely earliest Hettangian.Blue Lias FormationUnited KingdomA rhomaleosaurid. The type species is Avalonnectes arturi.thumb|150px|center|Avalonnectes.DjupedaliaEspen M. Knutsen Patrick S. Druckenmiller Jørn H. Hurum 2012 A new plesiosauroid (Reptilia: Sauropterygia) from the Agardhfjellet Formation (Middle Volgian) of central Spitsbergen, Norway Norwegian Journal of Geology 92 2–3 213–234 http://www.geologi.no/norwegian-journal-of-geology-91/details/1/56-56 Gen. et sp. nov.ValidKnutsen, Druckenmiller & HurumLate JurassicAgardhfjellet FormationNorwayA long-necked plesiosaurian. The type species is Djupedalia engeri.Dolichorhynchops tropicensisRebecca Schmeisser McKean 2012 A new species of polycotylid plesiosaur (Reptilia: Sauropterygia) from the Lower Turonian of Utah: extending the stratigraphic range of Dolichorhynchops Cretaceous Research 34 184–199 10.1016/j.cretres.2011.10.017 2012CrRes..34..184S Sp. nov.ValidMcKeanEarly TuronianTropic ShaleUnited StatesOriginally described as a third species of Dolichorhynchops, Clark, O'Keefe & Slack (2023) later moved it to the distinct genus Scalamagnus.Clark Robert O. O'Keefe F. Robin Slack Sara E. 2023-12-24 A new genus of small polycotylid plesiosaur from the Upper Cretaceous of the Western Interior Seaway and a clarification of the genus Dolichorhynchops https://www.sciencedirect.com/science/article/pii/S0195667123003403 Cretaceous Research 157 105812 10.1016/j.cretres.2023.105812 266546582 0195-6671 subscription EoplesiosaurusGen. et sp. novValidBenson, Evans & DruckenmillerEarly Jurassic, most likely earliest Hettangian.United KingdomA basal plesiosauroid. The type species is Eoplesiosaurus antiquior.thumb|150px|center|Eoplesiosaurus.HastanectesRoger B. J. Benson Hilary F. Ketchum Darren Naish Langan E. Turner 2013 A new leptocleidid (Sauropterygia, Plesiosauria) from the Vectis Formation (Early Barremian–early Aptian; Early Cretaceous) of the Isle of Wight and the evolution of Leptocleididae, a controversial clade Journal of Systematic Palaeontology 11 2 233–250 10.1080/14772019.2011.634444 2013JSPal..11..233B 18562271 Gen. et comb. novValid http://zoobank.org/References/8A0A8BD0-F6BB-4F74-BB18-79CBBA2F105C ZooBank.orgBenson et al.Early CretaceousUnited KingdomA new genus for "Cimoliasaurus" valdensisLusonectesAdam S. Smith Ricardo Araújo Octávio Mateus 2012 Lusonectes sauvagei, a new plesiosauroid from the Toarcian (Lower Jurassic) of Alhadas, Portugal Acta Palaeontologica Polonica 57 2 257–266 10.4202/app.2011.0023 55764533 free Gen. et sp. novValidSmith, Araújo & MateusToarcianSão Gião FormationPortugalA plesiosaurid plesiosaur.Pliosaurus funkeiEspen M. Knutsen Patrick S. Druckenmiller Jørn H. Hurum 2012 A new species of Pliosaurus (Sauropterygia: Plesiosauria) from the Middle Volgian of central Spitsbergen, Norway Norwegian Journal of Geology 92 2–3 235–258 http://www.geologi.no/norwegian-journal-of-geology-91/details/1/57-57 Sp. novValidKnutsen, Druckenmiller & HurumLate JurassicAgardhfjellet FormationNorwayA species of Pliosaurus.QianxisaurusYen-Nien Cheng Xiao-Chun Wu Tamaki Sato Hsi-Yin Shan 2012 A new eosauropterygian (Diapsida, Sauropterygia) from the Triassic of China Journal of Vertebrate Paleontology 32 6 1335–1349 10.1080/02724634.2012.695983 2012JVPal..32.1335C 85253142 Gen. et sp. novValidCheng et al.Middle Triassic (Ladinian)Falang FormationChinaAn eosauropterygian, a relative of pachypleurosaurs and nothosauroids. The type species is Qianxisaurus chajiangensis.SpitrasaurusEspen M. Knutsen Patrick S. Druckenmiller Jørn H. Hurum 2012 Two new species of long-necked plesiosaurians (Reptilia: Sauropterygia) from the Upper Jurassic (Middle Volgian) Agardhfjellet Formation of central Spitsbergen Norwegian Journal of Geology 92 2–3 187–212 http://www.geologi.no/norwegian-journal-of-geology-91/details/1/54-54 Gen. et 2 sp. nov.ValidKnutsen, Druckenmiller & HurumLate JurassicAgardhfjellet FormationNorwayA long-necked plesiosaurian. Genus contains two species: Spitrasaurus wensaasi and S. larseni.StratesaurusGen. et sp.ValidBenson, Evans & DruckenmillerEarly Jurassic, most likely earliest Hettangian.Blue Lias FormationUnited KingdomA rhomaleosaurid. The type species is Stratesaurus taylorithumb|150px|center|Stratesaurus.VectocleidusGen. et sp. novValidBenson et al.Early Cretaceous (late Barremian)Vectis FormationUnited KingdomA leptocleidid. The type species is Vectocleidus pastorum. Newly named rhynchocephalians Name Novelty Status Authors Age Unit Location Notes ImagesOenosaurusOliver W. M. Rauhut Alexander M. Heyng Adriana López-Arbarello Andreas Hecker 2012 A New Rhynchocephalian from the Late Jurassic of Germany with a Dentition That Is Unique amongst Tetrapods PLOS ONE 7 10 e46839 10.1371/journal.pone.0046839 23118861 3485277 2012PLoSO...746839R free Gen. et sp. novValidRauhut et al.Late Jurassic (early Tithonian)Mörnsheim FormationGermanyA sphenodontid rhynchocephalian. The type species is Oenosaurus muehlheimensis.SphenocondorSebastián Apesteguía Raúl O. Gómez Guillermo W. Rougier 2012 A basal sphenodontian (Lepidosauria) from the Jurassic of Patagonia: new insights on the phylogeny and biogeography of Gondwanan rhynchocephalians Zoological Journal of the Linnean Society 166 2 342–360 10.1111/j.1096-3642.2012.00837.x free 20.500.12110/paper_00244082_v166_n2_p342_Apesteguia free Gen. et sp. novValidApesteguía, Gómez & RougierMiddle JurassicArgentinaA rhynchocephalian lepidosaur. The type species is Sphenocondor gracilis. Squamates Research A large phylogenetic analysis of living and fossil squamates is published by Jacques A. Gauthier et al. (2012).Jacques A. Gauthier Maureen Kearney Jessica Anderson Maisano Olivier Rieppel Adam D.B. Behlke Assembling the Squamate Tree of Life: Perspectives from the Phenotype and the Fossil Record Bulletin of the Peabody Museum of Natural History 53 1 3–308 2012 10.3374/014.053.0101 2012BPMNH..53....3G 86355757 A study of squamate diversity in North America during the latest Cretaceous (Maastrichtian) and the impact of Cretaceous–Paleogene extinction event on the diversity of the group is published by Nicholas R. Longrich, Bhart-Anjan S. Bhullar and Jacques A. Gauthier (2012).Nicholas R. Longrich Bhart-Anjan S. Bhullar Jacques A. Gauthier 2012 Mass extinction of lizards and snakes at the Cretaceous–Paleogene boundary Proceedings of the National Academy of Sciences of the United States of America 109 52 21396–21401 10.1073/pnas.1211526110 23236177 3535637 2012PNAS..10921396L free New taxa Name Novelty Status Authors Age Unit Location Notes ImagesChianghsiaJin-you Mo Xing Xu Susan E. Evans 2012 A large predatory lizard (Platynota, Squamata) from the Late Cretaceous of South China Journal of Systematic Palaeontology 10 2 333–339 10.1080/14772019.2011.588254 2012JSPal..10..333M 85682211 Gen. et sp. novValidMo, Xu & EvansLate CretaceousNanxiong FormationChinaA platynotan lizard, probably a monstersaurian. The type species is Chianghsia nankangensis.DornosaurusV. R. Alifanov 2012 Lizards of the family Arretosauridae Gilmore, 1943 (Iguanomorpha, Iguania) from the Paleogene of Mongolia Paleontological Journal 46 4 412–420 10.1134/S0031030112040028 2012PalJ...46..412A 119087759 Gen. et sp. novValidAlifanovEoceneMongoliaAn arretosaurid iguanian. The type species is Dornosaurus gobiensis.EremiasaurusLeblanc A.R.H. Caldwell, M.W. Bardet, N. 2012 A new mosasaurine from the Maastrichtian (Upper Cretaceous) phosphates of Morocco and its implications for mosasaurine systematics Journal of Vertebrate Paleontology 32 1 82–104 10.1080/02724634.2012.624145 2012JVPal..32...82L 130559113 Gen. et sp. novValidLeblanc, Caldwell & BardetMaastrichtianMoroccoA mosasaur.ErgiliinsaurusGen. et sp. novValidAlifanovOligoceneErgiliin Zoo FormationMongoliaAn arretosaurid iguanian. The type species is Ergiliinsaurus postumus.JucarasepsBoler Arnau Susan E. Evans 2012 A tiny lizard (Lepidosauria, Squamata) from the Lower Cretaceous of Spain Palaeontology 55 3 491–500 10.1111/j.1475-4983.2012.01145.x 2012Palgy..55..491B 83645014 free Gen. et sp. novValidBolet & EvansEarly CretaceousSpainA lizard related to scleroglossan lizards. The type species is Jucaraseps grandipes.KhaichinguanaGen. et sp. novValidAlifanovEoceneKhaychin FormationMongoliaAn arretosaurid iguanian. The type species is Khaichinguana eocaenica.LouisamphisbaenaMarc Louis Augé 2012 Amphisbaenians from the European Eocene: a biogeographical review Palaeobiodiversity and Palaeoenvironments 92 4 425–443 10.1007/s12549-012-0104-6 2012PdPe...92..425A 129023376 Gen. et sp. novValidAugéEocene (Bartonian)FranceAn amphisbaenian lizard, possibly a member of Blanidae. The type species is Louisamphisbaena ferox.PannoniasaurusLászló Makádi Michael W. Caldwell Attila Ősi 2012 The First Freshwater Mosasauroid (Upper Cretaceous, Hungary) and a New Clade of Basal Mosasauroids PLOS ONE 7 12 e51781 10.1371/journal.pone.0051781 23284766 3526648 2012PLoSO...751781M free Gen. et sp. novValidMakádi, Caldwell & ŐsiLate Cretaceous (Santonian)Ajka Coal Formation Csehbánya FormationHungaryA tethysaurine mosasauroid closely related to Tethysaurus, Russellosaurus and Yaguarasaurus. The type species is Pannoniasaurus inexpectatus.?Placosaurus rageiRobert M. Sullivan Marc Augé Eric Wille Richard Smith 2012 A new glyptosaurine lizard from the earliest Eocene of Dormaal, Belgium Bulletin de la Société Géologique de France 183 6 627–633 10.2113/gssgfbull.183.6.627 Sp. nov.ValidSullivan et al.Earliest EoceneBelgiumA glyptosaurine anguid lizard. Originally described as a possible species of Placosaurus; Čerňanský et al. (2024) transferred it to the genus Gaultia.Andrej Čerňanský Richard Smith Thierry Smith Annelise Folie 2024 Timing of intercontinental faunal migrations: Anguimorph lizards from the earliest Eocene (MP 7) of Dormaal, Belgium Zoological Journal of the Linnean Society 201 4 zlae082 10.1093/zoolinnean/zlae082 PseudomimeosaurusV. R. Alifanov 2012 Отряд Lacertilia E.N. Kurochkin A. V. Lopatin Ископаемые позвоночные России и сопредельных стран. Ископаемые рептилии и птицы. Часть 2 / Fossil vertebrates of Russia and adjacent countries. Fossil Reptiles and Birds. Part 2 GEOS 7–136 978-5-89118-594-4 https://www.geokniga.org/books/11524 Gen. et comb. novValidAlifanovLate CretaceousDjadochta FormationMongoliaA pleurodontagamid iguanian lizard; a new genus for "Mimeosaurus" tugrikinensis Alifanov (1989).Pseudopus ahnikoviensisJozef Klembara 2012 A new species of Pseudopus (Squamata, Anguidae) from the early Miocene of Northwest Bohemia (Czech Republic) Journal of Vertebrate Paleontology 32 4 854–866 10.1080/02724634.2012.670177 2012JVPal..32..854K 129447096 Sp. novValidKlembaraEarly MioceneCzech RepublicAn anguid, a species of Pseudopus.PurbicellaSusan E. Evans Marc E. H. Jones Ryoko Matsumoto 2012 A new lizard skull from the Purbeck Limestone Group (Lower Cretaceous) of England Bulletin de la Société Géologique de France 183 6 517–524 10.2113/gssgfbull.183.6.517Gen. et sp. nov.ValidEvans, Jones & MatsumotoEarly CretaceousPurbeck Limestone GroupUnited KingdomA member of Lacertoidea. The type species isPurbicella ragei.RagesaurusSalvador Bailon Marc Auge 2012 A news genus, Ragesaurus (Squamata, Anguidae, Anguinae) from the lower Pleistocene of the islas Medas (Catalonia, Spain) Bulletin de la Société Géologique de France 183 6 683–688 10.2113/gssgfbull.183.6.683 Gen. et sp. nov.ValidBailon & AugeEarly PleistoceneSpainAn anguid lizard. The type species is Ragesaurus medasensis.Sphaerodactylus ciguapaJuan D. Daza Aaron M. Bauer 2012 A new amber-embedded sphaerodactyl gecko from Hispaniola, with comments on morphological synapomorphies of the Sphaerodactylidae http://www.mcz.harvard.edu/Publications/pubs/Breviora_2012_0529_2507.pdf Breviora 529 1–28 10.3099/529.1 34254367 Sp. novValidDaza & BauerLate Early Miocene or early Middle Miocene (20 to 15 MYA)Dominican RepublicA sphaerodactylid gecko found in Dominican amber, a species of Sphaerodactylus.Telmasaurus bialynickaeSp. novValidAlifanovLate CretaceousBarun Goyot FormationMongoliaA varanoid lizard, a species of Telmasaurus.UquiasaurusJuan D. Daza Virginia Abdala J. Salvador Arias Daniel García-López Pablo Ortiz 2012 Cladistic Analysis of Iguania and a Fossil Lizard from the Late Pliocene of Northwestern Argentina Journal of Herpetology 46 1 104–119 10.1670/10-112 11336/61054 85405843 free Gen. et sp. novDisputedDaza et al.Late PlioceneUquía FormationArgentinaAn iguanian lizard. The type species is Uquiasaurus heptanodonta. Scanferla & Díaz-Fernández (2023) reinterpreted the type series of this species as a fossil bone assemblage composed by more than one species of Liolaemus.Agustín Scanferla Linda Díaz-Fernánde. Reassessment of Uquiasaurus heptanodonta, an iguanian lizard from the late Pliocene (Uquía Formation) of NW Argentina 2023 Ameghiniana 10.5710/AMGH.01.09.2023.3577 261523182 Varanus (Varaneades) amnhophilisJack L. Conrad Ana M. Balcarcel Carl M. Mehling 2012 Earliest Example of a Giant Monitor Lizard (Varanus, Varanidae, Squamata) PLOS ONE 7 8 e41767 10.1371/journal.pone.0041767 22900001 3416840 2012PLoSO...741767C free Subgen. et sp. novDisputedConrad, Balcarcel & MehlingMiocene (Turolian)Mytilini FormationGreeceA monitor lizard. Villa et al. (2018) considered this species to be likely junior synonym of Varanus marathonensis.Andrea Villa Juan Abella David M. Alba Sergio Almécija Arnau Bolet George D. Koufos Fabien Knoll Àngel H. Luján Jorge Morales Josep M. Robles Israel M. Sánchez Massimo Delfino 2018 Revision of Varanus marathonensis (Squamata, Varanidae) based on historical and new material: morphology, systematics, and paleobiogeography of the European monitor lizards PLOS ONE 13 12 e0207719 10.1371/journal.pone.0207719 30517172 6281198 2018PLoSO..1307719V free Turtles Research A large phylogenetic analysis of basal turtles is published by Jérémy Anquetin (2012).Jérémy Anquetin 2012 Reassessment of the phylogenetic interrelationships of basal turtles (Testudinata) Journal of Systematic Palaeontology 10 1 3–45 10.1080/14772019.2011.558928 2012JSPal..10....3A 85295987 http://osf.io/dt6yj/ New taxa Name Novelty Status Authors Age Unit Location Notes ImagesAllopleuron qazaqstanenseHans-Volker Karl Elke Gröning Carsten Brauckmann 2012 New materials of the giant sea turtle Allopleuron (Testudines: Chelonioidea) from the marine Late Cretaceous of Central Europe and the Palaeogene of Kazakhstan Stvdia Palaeocheloniologica IV, Stvdia Geologica Salmanticensia Especial 9 153–173 http://campus.usal.es/~revistas_trabajo/index.php/Palaeocheloniologica/article/view/9224/9484Sp. novValidKarl, Gröning & BrauckmannEocene (Lutetian)KazakhstanA member of Cheloniidae, a species of Allopleuron.Axestemys cerevisiaNatasha S. Vitek 2012 Giant fossil soft-shelled turtles of North America Palaeontologia Electronica 15 1 13A http://palaeo-electronica.org/content/2012-issue-1-articles/210-giant-soft-shelled-turtles sp novValidVitekEoceneBridger FormationUnited StatesA trionychid, a species of Axestemys.Axestemys montinsanasp novValidVitekPaleoceneDenver Formation Fort Union Formation Melville FormationUnited StatesA trionychid, a species of Axestemys.BallerstedtiaHans-Volker Karl Elke Gröning Carsten Brauckmann Mike Reich 2012 Ballerstedtia bueckebergensis, a new turtle from the Early Cretaceous Wealden Facies of Germany (Testudines: Pleurosternidae) Stvdia Palaeocheloniologica IV, Stvdia Geologica Salmanticensia Especial 9 47–60 http://campus.usal.es/~revistas_trabajo/index.php/Palaeocheloniologica/article/view/9215/9475Gen. et comb. et sp. novValidKarl et al.Early CretaceousGermany United KingdomA member of Paracryptodira, a new genus for "Pleurosternon" typocardium Seeley (1869). Genus also contains a new species Ballerstedtia bueckebergensis.BashuchelysHaiyan Tong Igor Danilov Yong Ye Hui Ouyang Guangzhao Peng 2012 Middle Jurassic turtles from the Sichuan Basin, China: a review Geological Magazine 149 4 675–695 10.1017/S0016756811000859 2012GeoM..149..675T 129039148 Gen. et comb. et sp. novValidTong, Danilov, Ye, Ouyang & PengMiddle JurassicXiashaximiao FormationChinaA bashuchelyid cryptodiran, a new genus for "Chengyuchelys" zigongensis (Ye, 1982). Genus also contains a new species Bashuchelys youngi.BerruchelusAdán Pérez-García 2012 Berruchelus russelli, gen. et sp. nov., a paracryptodiran turtle from the Cenozoic of Europe Journal of Vertebrate Paleontology 32 3 545–556 10.1080/02724634.2012.658933 2012JVPal..32..545P 128616842 Gen. et sp. novValidPérez-GarcíaUpper PaleoceneFranceA paracryptodiran closely related to Compsemys. The type species is Berruchelus russelli.CarbonemysEdwin A. Cadena Daniel T. Ksepka Carlos A. Jaramillo Jonathan I. Bloch 2012 New pelomedusoid turtles from the late Palaeocene Cerrejón Formation of Colombia and their implications for phylogeny and body size evolution Journal of Systematic Palaeontology 10 2 313–331 10.1080/14772019.2011.569031 2012JSPal..10..313C 59406495 Gen. et sp. novValidCadena et al.Late PalaeoceneCerrejón FormationColombiaA podocnemidid turtle. The type species is Carbonemys cofrinii.ChuannanchelysGen. et comb. novValidTong, Danilov, Ye, Ouyang & PengMiddle JurassicXiashaximiao FormationChinaA bashuchelyid cryptodiran, a new genus for "Chengyuchelys" dashanpuensis (Fang, 1987).Foxemys trabantiMárton Rabi Haiyan Tong Gábor Botfalvai 2012 A new species of the side-necked turtle Foxemys (Pelomedusoides: Bothremydidae) from the Late Cretaceous of Hungary and the historical biogeography of the Bothremydini Geological Magazine 149 4 662–674 10.1017/S0016756811000756 2012GeoM..149..662R 128635040 sp novValidRabi, Tong & BotfalvaiSantonianCsehbánya FormationHungaryA bothremydid, a species of Foxemys.GalvecheloneAdán Pérez-García Xabier Murelaga 2012 Galvechelone lopezmartinezae gen. et sp. nov., a new cryptodiran turtle in the Lower Cretaceous of Europe Palaeontology 55 5 937–944 10.1111/j.1475-4983.2012.01154.x 2012Palgy..55..937P 128636057 free Gen. et sp. novValidPérez-García & MurelagaEarly CretaceousSpainA cryptodiran turtle. The type species is Galvechelone lopezmartinezae.GuangdongemysJulien Claude Jiang-Yong Zhang Jian-Jun Li Jin-You Mo Xue-Wen Kuang Haiyan Tong 2012 Geoemydid turtles from the Late Eocene Maoming basin, southern China Bulletin de la Société Géologique de France 183 6 641–651 10.2113/gssgfbull.183.6.641 Gen. et sp. nov.ValidClaude et al.Late EoceneChinaA geoemydid turtle. The type species is Guangdongemys pingi.HoyasemysAdán Pérez-García Marcelo S. de la Fuente Francisco Ortega 2012 Hoyasemys jimenezi gen. et sp. nov., a freshwater basal eucryptodiran turtle from the Lower Cretaceous of Spain Acta Palaeontologica Polonica 57 2 285–298 10.4202/app.2011.0031 54218702 free Gen. et sp. novValidPérez-García, Fuente & OrtegaLate BarremianCalizas de La Huérguina FormationSpainA basal eucryptodiran.IberoccitanemysAdán Pérez-García Francisco Ortega Xabier Murelaga 2012 A new genus of Bothremydidae (Chelonii, Pleurodira) in the Cretaceous of Southwestern Europe Geobios 45 2 219–229 10.1016/j.geobios.2011.03.001 2012Geobi..45..219P Gen. et comb. novValidPérez-García, Ortega & MurelagaLate Campanian to MaastrichtianFrance SpainA bothremydid, a new genus for "Elochelys" convenarum (Laurent et al. 2002). Pérez-García, Ortega & Murelaga (2021) considered I. convenarum to be a junior synonym of "Polysternon" atlanticum, resulting in a new combination Iberoccitanemys atlanticum.Adán Pérez-García Francisco Ortega Xabier Murelaga 2021 Iberoccitanemys atlanticum (Lapparent de Broin & Murelaga, 1996) n. comb.: new data on the diversity and paleobiogeographic distributions of the Campanian-Maastrichtian bothremydid turtles of Europe Comptes Rendus Palevol 20 32 667–676 10.5852/cr-palevol2021v20a32 238704135 free Kinosternon pojoaqueJason R. Bourque 2012 An extinct mud turtle of the Kinosternon flavescens group (Testudines, Kinosternidae) from the middle Miocene (late Barstovian) of New Mexico Journal of Vertebrate Paleontology 32 1 68–81 10.1080/02724634.2012.626824 2012JVPal..32...68B 84889382 SpeciesValidBourqueMiddle MioceneUnited StatesA species of Kinosternon.Kinosternon skullridgescensJason R. Bourque 2012 A fossil mud turtle (Testudines, Kinosternidae) from the early middle Miocene (early Barstovian) of New Mexico Journal of Vertebrate Paleontology 32 4 836–853 10.1080/02724634.2012.670179 2012JVPal..32..836B 85651690 SpeciesValidBourqueEarly middle MioceneTesuque FormationUnited StatesA species of Kinosternon.LarachelusAdán Pérez-García Xabier Murelaga 2012 Larachelus morla, gen. et sp. nov., a new member of the little-known European Early Cretaceous record of stem cryptodiran turtles Journal of Vertebrate Paleontology 32 6 1293–1302 10.1080/02724634.2012.694592 2012JVPal..32.1293P 84274421 Gen. et sp. novValidPérez-García & MurelagaEarly Cretaceous (late Hauterivian–early Barremian)Pinilla de los Moros FormationSpainA basal member of Pan-Cryptodira (the clade containing living cryptodirans and all extinct turtles that were more closely related to them than to pleurodirans). The type species is Larachelus morla.OerteliaHans-Volker Karl Stefanie Biermann Gottfried Tichy 2012 The oldest true sea turtle of the world, Oertelia gigantea (Oertel, 1914) n. gen. from the Aptian of Kastendamm near Hanover, Germany Stvdia Palaeocheloniologica IV, Stvdia Geologica Salmanticensia Especial 9 107–128 http://campus.usal.es/~revistas_trabajo/index.php/Palaeocheloniologica/article/view/9219/9479Gen. et comb. novValidKarl, Biermann & TichyEarly Cretaceous (early Aptian)GermanyA toxochelyid sea turtle, a new genus for "Toxochelys" gigantea Oertel (1914).PeligrochelysJuliana Sterli Marcelo S. de la Fuente 2013 New evidence from the Palaeocene of Patagonia (Argentina) on the evolution and palaeo-biogeography of Meiolaniformes (Testudinata, new taxon name) Journal of Systematic Palaeontology 11 7 835–852 10.1080/14772019.2012.708674 2013JSPal..11..835S 83804365 11336/3254 free Gen. et sp. novValid http://zoobank.org/References/C52A01FB-CC2B-4B9B-9B40-096CCCB9749A ZooBank.orgSterli & de la FuentePalaeoceneSalamanca FormationArgentinaA relative of meiolaniids. The type species is Peligrochelys walshae.Polysternon isonaeJ. Marmi Á.H. Luján V. Riera R. Gaete O. Oms À. Galobart 2012 The youngest species of Polysternon: A new bothremydid turtle from the uppermost Maastrichtian of the southern Pyrenees Cretaceous Research 35 133–142 10.1016/j.cretres.2011.12.004 2012CrRes..35..133M SpeciesValidMarmi et al..Late MaastrichtianSpainA bothremydid, a species of Polysternon.ProtoxinjiangchelysGen. et sp. novValidTong, Danilov, Ye, Ouyang & PengMiddle JurassicXiashaximiao FormationChinaA xinjiangchelyid cryptodiran.PuentemysEdwin A. Cadena Jonathan I. Bloch Carlos A. Jaramillo 2012 New Bothremydid Turtle (Testudines, Pleurodira) from the Paleocene of Northeastern Colombia Journal of Paleontology 86 4 688–698 10.1666/11-128R1.1 2012JPal...86..688C 85602476 Gen. et sp. nov.ValidCadena, Bloch & JaramilloPaleoceneCerrejón FormationColombiaA bothremydid turtle. The type species is Puentemys mushaisaensis.Rhinoclemmys panamaensisEdwin Cadena Jason R. Bourque Aldo F. Rincon Jonathan I. Bloch Carlos A. Jaramillo Bruce J. Macfadden 2012 New Turtles (Chelonia) from the Late Eocene Through Late Miocene of the Panama Canal Basin Journal of Paleontology 86 3 539–557 10.1666/11-106.1 2012JPal...86..539C 130656574 Sp novValidCadena et al.MioceneCucaracha FormationPanamaA geoemydid turtle, a species of Rhinoclemmys.Staurotypus moschusSp novValidCadena et al.MioceneCucaracha FormationPanamaA kinosternid turtle, a species of Staurotypus.Tasbacka danicaHans-Volker Karl Henrik Madsen 2012 Tasbacka danica n. sp., a new Eocene marine turtle of Denmark (Testudines: Chelonioidea) Stvdia Palaeocheloniologica IV, Stvdia Geologica Salmanticensia Especial 9 193–204 http://campus.usal.es/~revistas_trabajo/index.php/Palaeocheloniologica/article/view/9226/9486Sp. novValidKarl & MadsenEarly EoceneFur FormationDenmarkA member of Cheloniidae, a species of Tasbacka.Terrapene parornataWalter G. Joyce Andrea Petričević Tyler R. Lyson Nicholas J. Czaplewski 2012 A New Box Turtle from the Miocene/Pliocene Boundary (Latest Hemphillian) of Oklahoma and A Refined Chronology of Box Turtle Diversification Journal of Paleontology 86 1 177–190 http://jpaleontol.geoscienceworld.org/content/86/1/177.abstract 10.1666/11-073.1 2012JPal...86..177J 128857678 subscription Sp novValidJoyce et al..Miocene/Pliocene boundaryUnited StatesA box turtle.YuchelysHaiyan Tong, Li Xu Eric Buffetaut Xingliao Zhang Songhai Jia 2012 A new nanhsiungchelyid turtle from the Late Cretaceous of Neixiang, Henan Province, China Annales de Paléontologie 98 4 303–314 10.1016/j.annpal.2012.08.001 2012AnPal..98..303T Gen. et sp. novValidTong et al.Late CretaceousGaogou FormationChinaA nanhsiungchelyid cryptodiran. The type species is Yuchelys nanyangensis. Archosauromorphs Newly named basal archosauromorphs Name Novelty Status Authors Age Unit Location Notes ImagesChanaresuchus ischigualastensisMaría J. Trotteyn Ricardo N. Martínez Oscar A. Alcober 2012 A new proterochampsid Chanaresuchus ischigualastensis (Diapsida, Archosauriformes) in the early Late Triassic Ischigualasto Formation, Argentina Journal of Vertebrate Paleontology 32 2 485–489 10.1080/02724634.2012.645975 2012JVPal..32..485T 11336/199496 85962463 free sp novValidTrotteyn, Martínez & AlcoberLate TriassicIschigualasto FormationArgentinaA proterochampsian archosauriform. Originally described as a species of Chanaresuchus; subsequently made the type species of a separate genus Pseudochampsa by Trotteyn & Ezcurra (2014).María Jimena Trotteyn Martín D. Ezcurra 2014 Osteology of Pseudochampsa ischigualastensis gen. et comb. nov. (Archosauriformes: Proterochampsidae) from the Early Late Triassic Ischigualasto Formation of Northwestern Argentina PLOS ONE 9 11 e111388 10.1371/journal.pone.0111388 25426846 4245112 2014PLoSO...9k1388T free DiandongosuchusChun Li Xiao-Chun Wu Li-Jun Zhao Tamaki Sato Li-Ting Wang amp 2012 A new archosaur (Diapsida, Archosauriformes) from the marine Triassic of China Journal of Vertebrate Paleontology 32 5 1064–1081 10.1080/02724634.2012.694383 2012JVPal..32.1064L 86797826 Gen. et sp. nov.ValidLi et al.Middle Triassic (Ladinian)Falang FormationChinaA member of Archosauriformes of uncertain phylogenetic placement. Originally classified as a poposauroid pseudosuchian, but subsequently argued to be a phytosaur.Michelle R. Stocker Sterling J. Nesbitt Li-Jun Zhao Xiao-Chun Wu Chun Li 2016 Mosaic evolution in Phytosauria: the origin of long-snouted morphologies based on a complete skeleton of a phytosaur from the Middle Triassic of China Society of Vertebrate Paleontology 76th Annual Meeting Program & Abstracts 232 http://vertpaleo.org/PDFS/2016/SVP-2016-Program-Book-v10-with-covers.aspx 2016-10-30 2016-10-18 https://web.archive.org/web/20161018211820/http://vertpaleo.org/PDFS/2016/SVP-2016-Program-Book-v10-with-covers.aspx Michelle R. Stocker Li-Jun Zhao Sterling J. Nesbitt Xiao-Chun Wu Chun Li 2017 A Short-Snouted, Middle Triassic Phytosaur and its Implications for the Morphological Evolution and Biogeography of Phytosauria Scientific Reports 7 46028 10.1038/srep46028 free 28393843 5385495 2017NatSR...746028S The type species is Diandongosuchus fuyuanensis.thumb|150px|center|Diandongosuchus fuyuanensis.Doswellia sixmilensisAndrew B. Heckert Spencer G. Lucas Justin A. Spielmann 2012 A new species of the enigmatic archosauromorph Doswellia from the Upper Triassic Bluewater Creek Formation, New Mexico, USA Palaeontology 55 6 1333–1348 10.1111/j.1475-4983.2012.01200.x 2012Palgy..55.1333H 15431716 sp novValidHeckert, Lucas & SpielmannLate TriassicBluewater Creek FormationUnited StatesA doswelliid archosauromorph. Originally described as a species of Doswellia; Wynd et al. (2020) transferred it to the separate genus Rugarhynchos.Brenen M. Wynd Sterling J. Nesbitt Michelle R. Stocker Andrew B. Heckert 2020 A detailed description of Rugarhynchos sixmilensis, gen. et comb. nov. (Archosauriformes, Proterochampsia), and cranial convergence in snout elongation across stem and crown archosaurs Journal of Vertebrate Paleontology 39 6 e1748042 10.1080/02724634.2019.1748042 219917329 ProtomeMichelle R. Stocker 2012 A new phytosaur (Archosauriformes, Phytosauria) from the Lot's Wife beds (Sonsela Member) within the Chinle Formation (Upper Triassic) of Petrified Forest National Park, Arizona Journal of Vertebrate Paleontology 32 3 573–586 10.1080/02724634.2012.649815 2012JVPal..32..573S 129527672 Gen. et sp. novValidStockerLate TriassicChinle FormationUSAA phytosaur. The type species is Protome batalaria.150px Archosaurs 2012 in archosaur paleontology The following is a summary of the archosaurs described in 2012 8 pseudosuchians 1 basal dinosauriform 40 non-avian dinosaurs 64 birds 9 pterosaurs 1 archosaur of uncertain phylogenetic placement Synapsids Non-mammalian synapsids Research A phylogenetic analysis of basal synapsids is published by Roger B. J. Benson (2012).Roger B. J. Benson 2012 Interrelationships of basal synapsids: cranial and postcranial morphological partitions suggest different topologies Journal of Systematic Palaeontology 10 4 601–624 10.1080/14772019.2011.631042 2012JSPal..10..601B 84706899 New taxa Name Novelty Status Authors Age Unit Location Notes ImagesFortunodonA. A. Kurkin 2012 Dicynodontids of Eastern Europe Paleontological Journal 46 2 187–198 10.1134/S003103011201008X 2012PalJ...46..187K 84619486 Gen. et comb. novValidKurkinUpper PermianRussiaA dicynodont, a new genus for "Dicynodon" trautscholdi (Amalitzky, 1922) (considered to be a senior synonym of Vivaxosaurus permirus and assigned to the genus Vivaxosaurus by Kammerer et al., 2011).Kammerer C.F. Angielczyk, K.D. Fröbisch, J. 2011 A comprehensive taxonomic revision of Dicynodon (Therapsida, Anomodontia) and its implications for dicynodont phylogeny, biogeography, and biostratigraphy Journal of Vertebrate Paleontology 31 Suppl. 1 1–158 10.1080/02724634.2011.627074 2011JVPal..31S...1K 84987497 "Dicynodon" amalitzkii (Sushkin, 1926) is considered by Kurkin (2012) to be a second species of Fortunodon (Kammerer et al., 2011 classify it as the type species of the genus Peramodon.)NovocynodonM. F. Ivakhnenko 2012 Permian Cynodontia (Theromorpha) of Eastern Europe Paleontological Journal 46 2 199–207 10.1134/S0031030112020062 2012PalJ...46..199I 129223563 Gen. et sp. novValidIvakhnenkoMiddle PermianRussiaA thrinaxodontid cynodont. The type species is Novocynodon kutorgai.PampaphoneusJuan Carlos Cisneros Fernando Abdala Saniye Atayman-Güven Bruce S. Rubidge A. M. Celâl Şengör Cesar L. Schultz 2012 Carnivorous dinocephalian from the Middle Permian of Brazil and tetrapod dispersal in Pangaea Proceedings of the National Academy of Sciences of the United States of America 109 5 1584–1588 10.1073/pnas.1115975109 22307615 3277192 2012PNAS..109.1584C free Gen. et sp. novValidCisneros, Abdala, et al.Guadalupian (Middle Permian)Rio do Rasto FormationBrazilAn anteosaurid dinocephalian.PanchetocynodonD. P. Das Abir Gupta 2012 New cynodont record from the lower Triassic Panchet Formation, Damodar valley Journal of the Geological Society of India 79 2 175–180 10.1007/s12594-012-0022-2 2012JGSI...79..175D 129721800 Gen. et sp. novValidDas & GuptaEarly TriassicPanchet FormationIndiaA cynodont. The type species is Panchetocynodon damodarensis.PurbeckodonP. M. Butler D. Sigogneau-Russell P. C. Ensom 2012 Possible persistence of the morganucodontans in the Lower Cretaceous Purbeck Limestone Group (Dorset, England) Cretaceous Research 33 1 135–145 10.1016/j.cretres.2011.09.007 2012CrRes..33..135B Gen. et sp. novValidButler, Sigogneau-Russell, & EnsomEarly CretaceousPurbeck Limestone GroupUnited KingdomA possible morganucodontan.SludicaGen. et sp. novValidIvakhnenkoUpper PermianRussiaA procynosuchid cynodont. The type species is Sludica bulanovi. Mammals 2012 in mammal paleontology The following is a summary of the mammals described in 2012 9 non-eutherian mammals 24 eutherian mammals Other animals Name Novelty Status Authors Age Unit Location Notes ImagesAggerscolexJoseph P. Botting Lucy A. Muir Peter Van Roy Denis Bates Christopher Upton 2012 Diverse middle Ordovician palaeoscolecidan worms from the Builth-Llandrindod Inlier of central Wales Palaeontology 55 3 501–528 10.1111/j.1475-4983.2012.01135.x free 2012Palgy..55..501B Gen. et sp. novValidBotting et al.OrdovicianUnited KingdomA palaeoscolecid. The type species is A. murchisoni.Belebey augustodunensisJocelyn Falconnet 2012 First evidence of a bolosaurid parareptile in France (latest Carboniferous-earliest Permian of the Autun basin) and the spatiotemporal distribution of the Bolosauridae Bulletin de la Société Géologique de France 183 6 495–508 10.2113/gssgfbull.183.6.495Sp. nov.ValidFalconnetLatest Carboniferous (Gzhelian) or earliest Permian (Asselian)FranceAn amniote of uncertain phylogenetic placement. Originally classified as a bolosaurid and a species of Belebey; Spindler, Voigt & Fischer (2019) considered it to be a possible edaphosaurid synapsid instead.Frederik Spindler Sebastian Voigt Jan Fischer 2019 Edaphosauridae (Synapsida, Eupelycosauria) from Europe and their relationship to North American representatives PalZ 94 125–153 10.1007/s12542-019-00453-2 198140317 BullascolexGen. et sp. novValidBotting et al.OrdovicianUnited KingdomA palaeoscolecid. The type species is B. inserere.CarbotubulusJoachim T. Haug Georg Mayer Carolin Haug Derek E.G. Briggs 2012 A Carboniferous Non-Onychophoran Lobopodian Reveals Long-Term Survival of a Cambrian Morphotype Current Biology 22 18 1673–1675 10.1016/j.cub.2012.06.066 22885062 17371138 free 2012CBio...22.1673H Gen. et sp. novValidHaug et al.CarboniferousCarbondale FormationUnited StatesA long-legged lobopodian. The type species is C. waloszeki.Circotheca smetanaiMartin Valent Oldřich Fatka Michal Szabad Václav Micka Ladislav Marek 2012 Two new orthothecids from the Cambrian of the Barrandian area (Hyolitha, Skryje-Týřovice Basin, Czech Republic) Bulletin of Geosciences 87 2 241–248 10.3140/bull.geosci.1142 free Sp. novValidValent et al.Middle CambrianBuchava FormationCzech RepublicA member of Hyolitha (a group of animals of uncertain phylogenetic placement, possibly molluscs), a species of Circotheca.CoronacollinaErica C. Clites Mary L. Droser James G. Gehling 130837736 2012 The advent of hard-part structural support among the Ediacara biota: Ediacaran harbinger of a Cambrian mode of body construction Geology 40 4 307–310 10.1130/G32828.1 2012Geo....40..307C Gen. et sp. nov.ValidClites, Droser & GehlingEdiacaranAustraliaAn animal of uncertain phylogenetic placement. The type species is Coronacollina acula.Crenulipora kuwanoiShuji Niko 2012 Additional Material of Favositid and Auloporid Tabulate Corals from the Devonian Fukuji Formation, Gifu Prefecture, Japan Bulletin of the National Museum of Nature and Science, Series C 38 47–62 https://www.kahaku.go.jp/research/publication/geology/download/38/BNMNS_C38_47-62.pdf Sp. novValidNikoDevonianFukuji FormationJapanA tabulate coral belonging to the group Favositida and the family Favositidae.DispinoscolexBaichuan Duan Xi-Ping Dong Philip C. J. Donoghue 2012 New palaeoscolecid worms from the Furongian (upper Cambrian) of Hunan, South China: is Markuelia an embryonic palaeoscolecid? Palaeontology 55 3 613–622 10.1111/j.1475-4983.2012.01148.x 2012Palgy..55..613D https://www.dora.lib4ri.ch/psi/islandora/object/psi%3A11695 free subscription Gen. et sp. novValidDuan, Dong & DonoghueCambrian (Paibian)Bitiao FormationChinaA palaeoscolecid. The type species is D. decorus.Favia maitreyiaeRamón Andrés López-Pérez 2012 Late Miocene to Pleistocene Reef Corals in the Gulf of California Bulletins of American Paleontology 383 1–78 https://www.researchgate.net/publication/233181933 Sp. novValidLópez-PérezPlioceneCarmen FormationMexicoA stony coral, a species of Favia.Favia tulsidasiSp. novValidLópez-PérezPlioceneSan Marcos FormationMexicoA stony coral, a species of Favia.Heydenius arachniusGeorge O. Poinar Jr 86060924 2012 New fossil nematodes in Dominican and Baltic amber Nematology 14 4 483–488 10.1163/156854111X612199 Sp. novValidPoinarEocene to MioceneDominican amberDominican RepublicA nematode belonging to the family Mermithidae.Heydenius phasmatophilusSp. novValidPoinarEoceneBaltic amberRussia(Kaliningrad Oblast)A nematode belonging to the family Mermithidae.Heydenius podenasaeSp. novValidPoinarEoceneBaltic amberRussia(Kaliningrad Oblast)A nematode belonging to the family Mermithidae.Heydenius trichorosusSp. novValidPoinarEoceneBaltic amberRussia(Kaliningrad Oblast)A nematode belonging to the family Mermithidae.LoriciscolexGen. et sp. novValidBotting et al.OrdovicianUnited KingdomA palaeoscolecid. The type species is L. cuspidus.Parastriatopora bolivianaSp. novValidPlusquellec, Tourneur & Fernández-Martínez in Racheboeuf et al.Devonian (Emsian or Eifelian)Belén FormationBoliviaA tabulate coral belonging to the superfamily Pachyporoidea and the family Parastriatoporidae.Placosmilia? aliciaeSp. novValidLópez-PérezPlioceneSan Marcos Formation San Nicolas FormationMexicoA stony coral belonging to the family Montlivaltiidae.PluoscolexGen. et sp. novValidBotting et al.OrdovicianUnited KingdomA palaeoscolecid. The type species is P. linearis.ProbactrothecaGen. et sp. novValidValent et al.Middle CambrianBuchava FormationCzech RepublicA member of Hyolitha. The type species is Probactrotheca briketa.Protomichelinia foveataShuji Niko Takehiko Haikawa Masayuki Fujikawa 2012 Permian tabulate corals from the Akiyoshi Limestone Group, Yamaguchi Prefecture Bulletin of the Akiyoshi-dai Museum of Natural History 47 5–10 Sp. novValidNiko, Haikawa & FujikawaPermianAkiyoshi Limestone GroupJapanA tabulate coral.Pseudoromingeria murataiSp. novValidNiko, Haikawa & FujikawaPermianAkiyoshi Limestone GroupJapanA tabulate coral.RadnorscolexGen. et sp. novValidBotting et al.OrdovicianUnited KingdomA palaeoscolecid. The type species is R. bwlchi.RedondagnathusJustin A. Spielmann Spencer G. Lucas 2012 Tetrapod fauna of the Upper Triassic Redonda Formation, east-central New Mexico; the characteristic assemblage of the Apachean land-vertebrate faunachron New Mexico Museum of Natural History and Science Bulletin 55 1–119 Gen. et sp. novValidSpielmann & LucasLate TriassicUnited StatesA vertebrate of uncertain phylogenetic placement. Described on the basis of three postcanine teeth, considered to be teeth of a trirachodontid cynodont by Lucas et al. (1999)Spencer G. Lucas John W. Estep Andrew B. Heckert Adrian P. Hunt 1999 Cynodont teeth from the Upper Triassic of New Mexico, USA Neues Jahrbuch für Geologie und Paläontologie - Monatshefte Jg. 1999 6 331–344 and Spielmann and Lucas (2012). On the other hand, Abdala, Neveling and Welman (2006) and Sidor and Hopson (2018) did not consider it likely that Redondagnathus was a member of Trirachodontidae, and Abdala, Neveling and Welman (2006) weren't even sure whether it was a cynodont.F. Abdala J. Neveling J. Welman 2006 A new trirachodontid cynodont from the lower levels of the Burgersdorp Formation (Lower Triassic) of the Beaufort Group, South Africa and the cladistic relationships of Gondwanan gomphodonts Zoological Journal of the Linnean Society 147 3 383–413 10.1111/j.1096-3642.2006.00224.x free Christian A. Sidor James A. Hopson 2018 Cricodon metabolus (Cynodontia: Gomphodontia) from the Triassic Ntawere Formation of northeastern Zambia: patterns of tooth replacement and a systematic review of the Trirachodontidae Journal of Vertebrate Paleontology 37 Supplement to No. 6 39–64 10.1080/02724634.2017.1410485 89932366 The type species is Redondagnathus hunti.Schistoscolex hunanensisSp. novValidDuan, Dong & DonoghueCambrian (Paibian)Bitiao FormationChinaA palaeoscolecid.Siderastrea annaeSp. novValidLópez-PérezPlioceneSan Marcos FormationMexicoA stony coral, a species of Siderastrea.SinopathesAndrzej Baliński Yuanlin Sun Jerzy Dzik 2012 470-Million-year-old black corals from China Naturwissenschaften 99 8 645–653 10.1007/s00114-012-0947-8 22790835 3407557 2012NW.....99..645B Gen. et sp. novValidBaliński, Sun & DzikEarly OrdovicianFenxiang FormationChinaA black coral. The type species is S. reptans.SiphusauctumLorna J. O'Brien Jean-Bernard Caron 2012 A New Stalked Filter-Feeder from the Middle Cambrian Burgess Shale, British Columbia, Canada PLOS ONE 7 1 e29233 10.1371/journal.pone.0029233 3261148 22279532 2012PLoSO...729233O free Gen. et sp. novValidO'Brien & CaronMiddle CambrianBurgess ShaleCanadaA stem-bilaterian of uncertain phylogenetic placement, a stalked filter feeder. The type species is S. gregarium.Syringopora konishiiShuji Niko Yousuke Ibaraki 2012 Syringopora konishii, a new species of Early Carboniferous tabulate corals from the allochthonous limestone blocks in the Kotaki Formation, Niigata Prefecture, central Japan Science Reports of Niigata University. (Geology) 27 9–14 10191/22745 Sp. novValidNiko & IbarakiCarboniferous (Viséan)Kotaki FormationJapanA tabulate coral belonging to the group Auloporida.Thecia ichikawaiShuji Niko Tomio Adachi 2012 Silurian Favositids (Coelenterata: Tabulata) from the Gionyama Formation, Miyazaki Prefecture, Japan Bulletin of the National Museum of Nature and Science, Series C 38 33–46 https://www.kahaku.go.jp/research/publication/geology/download/38/BNMNS_C38_33-46.pdf Sp. novValidNiko & AdachiSilurianGionyama FormationJapanA tabulate coral belonging to the group Favositida and the family Theciidae.Trachypsammia konboShuji Niko 2012 Trachypsammia konbo, a new species of Early Carboniferous tabulate corals from the Akiyoshi Limestone Group, Yamaguchi Prefecture Bulletin of the Akiyoshi-dai Museum of Natural History 47 1–4 Sp. novValidNikoEarly CarboniferousAkiyoshi Limestone GroupJapanA tabulate coral.UlexiscolexGen. et sp. novValidBotting et al.OrdovicianUnited KingdomA palaeoscolecid. The type species is U. ormrodi.WerniaGen. et sp. novValidBotting et al.OrdovicianUnited KingdomA palaeoscolecid. The type species is W. eximia. Other organisms Name Novelty Status Authors Age Unit Location Notes ImagesHydrolithon braganumWilliam J. Woelkerling Davide Bassi Yasufumi Iryu 2012 Hydrolithon braganum sp. nov. (Corallinaceae, Rhodophyta), the first known exclusively fossil semi-endophytic coralline red alga Phycologia 51 6 604–611 10.2216/11-119.1 2012Phyco..51..604W 84505820 Sp. novValidWoelkerling, Bassi & IryuQuaternaryReef terraces around Tahiti, French PolynesiaA red alga belonging to the family Corallinaceae, a species of Hydrolithon.JimwhiteaMichael Krings Thomas N. Taylor Nora Dotzler Gianna Persichini 2012 Fossil fungi with suggested affinities to the Endogonaceae from the Middle Triassic of Antarctica Mycologia 104 4 835–844 10.3852/11-384 22453117 1808/13692 13391117 free Gen. et sp. novValidKrings & Taylor in Krings et al.Middle TriassicFremouw FormationAntarcticaA fungus belonging to the group Endogonales. The type species is Jimwhitea circumtecta. References 3 Category:2010s in paleontology Paleontology